Annual or perennial herbs, simple or branched, sometimes tufted; stems quadrangular, glabrous to glandular-villous or hirsute
Leaves
mostly opposite, sometimes alternate near inflorescence, petiolate, lanceolate to ovate, entire or dentate to pinnatifid, apex acute
Flowers
1-several in lax terminal racemes, alternate, resupinate by twisting of pedicels
Bracts
leafy or bractiform, ovate to linear
Calyx
5-lobed; segments lanceolate or narrowly elliptical, somewhat unequal, entire or irregularly incised, imbricate at base in bud
Corolla
expanded, bilabiate, unequally 5-lobed, tube very short, split to base on anterior side (morphologically posterior side); limb spreading; upper lip (actually lower one) 3-lobed, central one largest, broadly ovate, lateral ones ovate or broadly so, gamopetalous portion at base of lateral lobes with 2 shallow sacs lined with oil-secreting hairs; lower lip (actually upper one) split centrally to base, of 2 broad lobes, exterior in bud
Stamens
4, all perfect, didynamous; filaments short, basally declinate, not twisted round each other, glabrous, or glandular-puberulent; anthers shortly oblong, unithecate by confluence; staminodes 0
Ovary
bilocular, ovate or widely ovate, obtuse or emarginate; ovules many; style involute, subulate, persistent; stigma capitate or minutely bifid
Fruit
an ovate or pyriform, emarginate, septicidally bivalved capsule; valves entire, emarginate or bifid
Seeds
many, small, ovoid or oblong to elliptic, wingless, a little depressed at both ends, dark brown with shallow longitudinal grooves, punctate-rugose or papillate
x = 7, 12, 25 (1 report each, aneuploids, polyploidy)
Nomenclature:
Alonsoa
Ruiz & Pav.
Ruiz & Pavón: 150 (1798)
Wettstein: 53 (1891/1893)
Hiern: 138 (1904)
Hilliard & Burtt: 338 (1984)
Roux: 7 (1986)
Steiner: 1152 (1989)
Steiner: 63 (1996)
Schistanthe
Kunze
Kunze: 109 (1842)
Bentham: 249 (1846)
Wettstein: 53 (1891/1893)
Distribution & Notes:
Global
: Species 16, mainly South American, ranging from Mexico to Chile, centred in Peru
Southern Africa
: Species 2:
Alonsoa peduncularis
(Kunze) Wettst., Western and Eastern Cape (restricted to Rooiberg, Touwsberg and near Uitenhage), and
A. unilabiata
(L.f.) Steud. from Western Cape, but probably extinct near Cape Town
South African species of
Alonsoa
differ from South American ones in having 2 shallow, trichome-lined sacs on corolla
Oil-collecting bees act as pollinators
References:
BENTHAM, G. 1846.
Scrophulariaceae
. In A.P. de Candolle,
Prodromus
10. Masson, Paris
HILLIARD, O.M. & BURTT, B.L. 1984. A revision of
Diascia
section
Racemosae
.
Journal of South African Botany
50
KUNZE, G. 1842.
Schistanthe peduncularis
Kunze.
Linnaea
16
ROUX, J.P. 1986.
Alonsoa peduncularis
rediscovered.
South African Journal of Botany
52
RUIZ, H. & PAVÓN, J.A. 1798.
Systema vegetabilium florae peruvianae et chilensis
. Gabrielis de Sancha (typ.), Madrid
STEINER, K.E. 1989. A second species of the amphi-Atlantic genus
Alonsoa
(
Scrophulariaceae
) in South Africa.
Annals of the Missouri Botanical Garden
76
STEINER, K.E. 1996. Chromosome numbers and relationships in tribe
Hemimerideae
(
Scrophulariaceae
).
Systematic Botany
21
WETTSTEIN, R. VON. 1891/1893.
Scrophulariaceae
.
Die natürlichen Pflanzenfamilien
4, 3b
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