e-Key v3 - Holcus
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Interactive keys to the identification of seed plants of southern Africa using keys based on plant morphology.

Poaceae - Pooideae - Aveneae - Aveninae - Holcus L.

Description :

  • Annual or perennial, tufted
  • Leaf blade linear, expanded, pubescent or pilose; ligule an unfringed or fringed membrane
  • Inflorescence a panicle, contracted, often dense, sometimes almost spike-like, oblong or interrupted; spikelets solitary, pedicelled, pedicel terete, pilose
  • Spikelet laterally compressed, falling with glumes; glumes ± equal, ± as long as to longer than spikelet, membranous, dissimilar, keeled; lower glume lanceolate, hairy on keels and margins, 1-3-nerved, awned; awn half as long as body of glume; upper glume larger, elliptic-lanceolate or ovate, 3-nerved, with 2 prominent lateral nerves, hairy on keel and margins, shortly mucronate or awned; awn as long as body of glume
  • Florets 2, a brief rachilla extension often present; lower floret bisexual; lemma firmer in texture than glumes, shiny, ovate, rounded on back, obscurely 3-5-nerved, obtuse or acute, awnless; upper floret specialised in form, male or sterile, lemma shorter to ± as long as lower lemma, lanceolate, hyaline or membranous, glabrous, awned from back near apex; awn short, stout, hooked, usually geniculate, slightly twisted; palea smaller than lemma, narrow-lanceolate
  • Lodicules 2, delicate
  • Stamens 3
  • Ovary glabrous; styles distinct, plumose above
  • Caryopsis narrowly ellipsoid
  • x = 4, 7 (polyploidy)

Nomenclature:

  • Holcus L.
    • Linnaeus: 1047 (1753) name conserved
    • Stapf: 464 (1899)
    • Stent: 284 (1924)
    • Chippindall: 87 (1955)
    • Tutin: 230 (1980)
    • Clayton & Renvoize: 130 (1986)
    • Gibbs Russell et al.: 180 (1990)
    • Watson & Dallwitz: 464 (1994)

Distribution & Notes:

  • Global : Species 6, Europe, North Africa and Middle East
  • Southern Africa : Species ± 2, 1 endemic: Holcus setiger Nees, 1 naturalised: * H. lanatus L., northern provinces of South Africa, KwaZulu-Natal, Eastern and Western Cape

References:

  • CHIPPINDALL, L.K.A. 1955. A guide to the identification of grasses in South Africa. In D. Meredith, The grasses and pastures of South Africa . Central News Agency, Cape Town
  • CLAYTON, W.D. & RENVOIZE S.A. 1986. Genera graminum. Grasses of the world. Kew Bulletin . Additional series 13
  • GIBBS RUSSELL, G.E., WATSON, L., KOEKEMOER, M., SMOOK, L., BARKER, N.P., ANDERSON, H.M. & DALLWITZ. M.J. 1990. Grasses of southern Africa. Memoirs of the Botanical Survey of South Africa No. 58
  • LINNAEUS, C. 1753. Species plantarum . Laurentius Salvius, Stockholm
  • STAPF, O. 1898-1900. Gramineae . Flora capensis 7
  • STENT, S.M. 1924. South African Gramineae . Grasses of the Transvaal as represented in the National Herbarium. Bothalia 1
  • TUTIN, T.G. 1980. Flora europaea 5
  • WATSON, L. & DALLWITZ, M.J. 1994. The grass genera of the world , revised edn. CAB International, Oxon