Decumbent to erect shrublets or subshrubs, often with thickened roots, or prostrate to decumbent geophytes; stems corky or lignified throughout or only basally so, epidermal bladder cells conspicuous, occasionally hair-like, rarely flattened
Leaves
decussate throughout or becoming alternate in inflorescences or alternate throughout, subcylindrical, rarely flattened, shortly fused towards bases, deciduous or drying and persistent (marcescent)
Flowers
in dichasia or solitary, (10-)20-30(-40) mm in diameter; in most species flowers close at night, some open at night and others never close
Sepals
4 or 5, often fused basally into a short tube
Petals
in various shades of green, yellow, orange, red, pink or whitish
Staminodes
present or 0
Nectary
consisting of 5 separate grooves
Ovary
mostly conical on top; placentas axile; stigmas 4 or 5, subulate
Fruit
a 4- or 5-locular capsule, of Mesembryanthemum type; expanding keels extending to centre of fruit; covering membranes 0; valve wings mostly present and inflexed over valves or rarely reflexed and fused in pairs or 0
Seeds
D-shaped or horseshoe-shaped, 1.1-1.8 mm long, usually black without a crest, or brown with a crest, surface rough, rarely smooth
Flowering
from mid-winter to mid-summer
Distinguishing characters
:
Perennials, with conspicuous bladder cell idioblasts on leaves
If rootstock is tuberous, leaves are summer-deciduous
Nomenclature:
Phyllobolus
N.E.Br.
Brown: 413 (1925)
Brown: 1 (1926)
Gerbaulet: 385 (1995)
Gerbaulet: 145 (1997)
Smith et al.: 42 (1998)
Amoebophyllum
N.E.Br.
Brown: 433 (1925)
Herre: 68 (1971)
Sphalmanthus
N.E.Br.
Brown: 433 (1925)
Friedrich: 126 (1970)
Herre: 290 (1971)
Distribution & Notes:
Southern Africa
: Species 31, widespread in S Namibia, and in the western and central parts of South Africa (Northern, Western, and Eastern Cape and Free State)
References:
BROWN, N.E. 1925.
Mesembryanthemum
and some new genera separated from it.
Gardeners' Chronicle
78
BROWN, N.E. 1926.
Phyllobolus
. In E.P. Phillips, The genera of South African flowering plants.
Memoirs of the Botanical Survey of South Africa
No. 25
FRIEDRICH, H.C. 1970.
Aizoaceae
.
Prodromus einer Flora von Südwestafrika
27
GERBAULET, M. 1995.
Phyllobolus
N.E.Br. emend. Bittrich (
Aizoaceae
): A reassessment of generic boundaries.
Botanische Jahrbücher
117
GERBAULET, M. 1997. Revision of the genus
Phyllobolus
N.E.Br. (
Aizoaceae
).
Botanische Jahrbücher
119
HERRE, H. 1971.
The genera of the
Mesembryanthemaceae
: 68, 290. Tafelberg, Cape Town
SMITH, G.F., CHESSELET, P., VAN JAARSVELD, E.J., HARTMANN, H., HAMMER, S., VAN WYK, B-.E., BURGOYNE, P., KLAK, C. & KURZWEIL, H. 1998.
Mesembs of the world
. Briza, Pretoria
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