Annual, biennial or perennial herbs, rarely geophytes or shrublets, sometimes caespitose; plants often monoecious or rarely gynodioecious or dioecious
Leaves
opposite, entire, sometimes slightly connate at base; stipules 0
Inflorescences
dichasial or monochasial, sometimes ± capitate, sometimes flowers solitary
Flowers
bisexual or unisexual; anthophore present
Calyx
: tube 10-30(-60)-nerved, sometimes strongly inflated, with 5 short lobes, often glandular
Petals
5, arising on margin of disc, with long claw and entire to deeply divided limb, often with 2 scales at base
Stamens
10, in male and bisexual flowers, sometimes abnormally reduced, arising on margin of disc, the 5 opposite petals sometimes attached to claw
Disc
columnar, often pubescent
Ovary
on long gynophore adnate to disc, usually 1-locular, sometimes 3-locular towards base; ovules many on free central placenta; styles (2-)5, usually 3, as long as or longer than ovary, often glandular
Fruit
a capsule, opening usually by twice as many teeth as styles present
Seeds
many, ± reniform, compressed, reticulate or tuberculate, sometimes with 2 wings; embryo ± annular
x = 12 (aneuploids, high polyploidy)
Nomenclature:
Silene
L.
Linnaeus: 416 (1753)
Linnaeus: 193 (1754)
Jussieu: 302 (1789)
Sonder: 125 (1860)
Pax: 70 (1889)
Burtt Davy: 149 (1926)
Pax & Hoffmann: 343 (1934)
Wild: 350 (1961)
Chater & Walters: 158 (1964)
Henderson & Anderson: 138 (1966)
Podlech: 4 (1967)
Bittrich: 233 (1993)
Melandrium
Mertens & Koch
Mertens & Koch: 274 (1812)
Adamson: 399 (1950)
Distribution & Notes:
Global
: Species ± 700, in Eurasia (± 600 species), Africa (± 92 species), North America (± 50 species)
Southern Africa
: Species ± 19, of which 3 are naturalised, widespread in all provinces and countries except Botswana
References:
ADAMSON, R.S. 1950.
Caryophyllaceae
. In R.S. Adamson & T.M. Salter,
Flora of the Cape Peninsula
. Juta, Cape Town
BITTRICH, V. 1993.
Caryophyllaceae
. In K. Kubitzki, J.G. Rohwer & V. Bittrich,
The families and genera of vascular plants - dicotyledons
2. Springer-Verlag, Berlin
BURTT DAVY, J. 1926.
Caryophyllaceae
.
A manual of the flowering plants and ferns of the Transvaal with Swaziland
1. Longmans, London
WILD, H. 1961.
Caryophyllaceae
.
Flora zambesiaca
1
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