e-Key v3 - Holubia
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Interactive keys to the identification of seed plants of southern Africa using keys based on plant morphology.

Pedaliaceae - Holubia Oliv.

Description :

  • Annual herb, sometimes biennial, erect with spreading branches; stems square, slightly fleshy and minutely glandular
  • Leaves opposite, long-petiolate, circular to ovate, slightly lobed or sinuate, basally truncate or slightly cordate, subsucculent, glabrous above, sparingly glandular below
  • Flowers solitary, axillary; yellow-green to white
  • Calyx much shorter than corolla tube; segments lanceolate, glandular
  • Corolla : tube with large, posterior, flattened sac at base, centrally cylindrical and funnel-shaped at mouth; limb spreading, obscurely 2-lipped; lobes transversely oblong
  • Stamens didynamous, arising about middle of corolla tube, included; filaments long, terete, glandular-villous at base; anthers with thecae divergent, connective apiculate; staminode 1
  • Nectary thick, cushion-like, slightly asymmetrical
  • Ovary 2-locular, 4-angled, glandular; locules undivided with ± 8 ovules in each, in 2 rows, ascending; style terete, scarcely exserted; stigma of 2 ovate, somewhat crenate lobes
  • Fruit indehiscent, rotund to circular in lateral view, ± rectangular in cross section, each edge with a broad parchment-like wing with conspicuous, usually dark, reticulate veins; mesocarp not spongy
  • Seeds flattened, obovate, with a narrow, ribbed, marginal wing; testa finely honeycombed

Nomenclature:

  • Holubia Oliv.
    • Oliver: t. 1475 (1884)
    • Stapf: 457 (1904)
    • Phillips: t. 661 (1937)
    • Ihlenfeldt: 43. t. 7 (1967)
    • Ihlenfeldt: 89 (1988)
    • Ihlenfeldt: 1063 (1994)

Distribution & Notes:

  • Global : Monotypic: Holubia saccata Oliv., southern Africa and Zimbabwe
  • Southern Africa : Northern Province to E North-West and N Mpumalanga

References:

  • IHLENFELDT, H.-D. 1967. Über die Abgrenzung und die natürliche Gliederung der Pedaliaceae R. Br. Mitteilungen aus dem Staatsinstitut für Allgemeine Botanik Hamburg 12
  • IHLENFELDT, H.-D. 1988. Pedaliaceae . Flora zambesiaca 8, 3
  • IHLENFELDT, H.-D. 1994. Phytogeography of Pedaliaceae R.Br. In J.H. Seyani & A.C. Chikuni, Proceedings of the XIIIth Plenary Meeting of AETFAT, Malawi 2
  • OLIVER, D. 1884. Holubia saccata Oliv. Hooker's Icones Plantarum ser. 3,5
  • PHILLIPS, E.P. 1937. Holubia saccata . The Flowering Plants of South Africa 17
  • STAPF, O. 1904. Pedalineæ . Flora capensis 4,2