Annual herbs, or rarely small, woody, perennial subshrubs (
S. dulcis
), erect, fairly robust, much-branched, glabrous or minutely pubescent, especially at nodes; stems slightly angular or ribbed because of decurrent leaf bases
Leaves
commonly ternate, whorled or opposite, narrowed to petiole-like base, narrowly oblanceolate to elliptic-lanceolate, entire or dentate in upper half, usually narrow, punctate, veins prominent below
Flowers
small, usually solitary or more rarely paired in axils of leaf-like bracts, sometimes forming branched, many-flowered, raceme-like inflorescences, pedicellate, ebracteolate
Calyx
deeply 4(5)-lobed; lobes equal or subequal, ovate or oblong, obtuse, imbricate, densely sessile-glandular
Corolla
regular, rotate, 4(5)-lobed, corolla tube very short, not exceeding calyx tube, throat bearded; lobes obtuse, subequal, bearded in lower part, upper lobes probably outside in bud
Stamens
4, subequal, exserted; filaments filiform; anthers subsagittate with 2 divergent or parallel thecae
Ovary
bilocular, ovoid; ovules many; style subclavate at apex, persistent; stigma truncate or emarginate
Fruit
a globose or ovoid, septicidal capsule, when partly opened forming a ± round pore at apex; valves ± entire, membranous
Seeds
many, obovoid, angular, minutely pitted
x = 10 (polyploidy)
Classification Notes:
The arrangement of corolla lobes in bud of *
Scoparia
is uncertain
Wettstein (1891/1893) and Hemsley & Skan (1906) grouped it with
Veronica
in
Digitaleae
, which has upper lobes inside in bud
Dyer: 540 (1975) placed it in
Selaginaceae
Mielcarek (1996) and Takhtajan: 457 (1997) put it in subfamily
Scrophularioideae
,
Gratioleae
, these taxa having upper lobes outside in bud
The available material precluded checking of this feature
Nomenclature:
*Scoparia
L.
Linnaeus: 116 (1753)
Hemsley & Skan: 354 (1906)
Philcox: 77 (1990)
Mielcarek: 32 (1996)
Distribution & Notes:
Global
: Species 20; tropical America
Southern Africa
: Species 1: *
Scoparia dulcis
L., a pantropical weed of cultivation, grazed grasslands and wasteland; sometimes an escape in KwaZulu-Natal (Richards Bay, Inanda and Verulam Districts)
References:
DYER, R.A. 1975.
The genera of southern African flowering plants
, Vol. 1. Botanical Research Institute, Pretoria
HEMSLEY, W.B. & SKAN, S.A. 1906.
Scrophulariaceae
.
Flora of tropical Africa
4
LINNAEUS, C. 1753.
Species plantarum
, edn 1. Laurentius Salvius, Stockholm
MIELCAREK, R. 1996. Les
Scrophulariaceae
dans la flore d'Afrique centrale (excl.
Linderneae
).
Fragmenta floristica et geobotanica
41
PHILCOX, D. 1990.
Scrophulariaceae
.
Flora zambesiaca
8,2
TAKHTAJAN, A. 1997.
Diversity and classification of flowering plants
. Columbia University Press, New York
WETTSTEIN, R. VON. 1891/1893.
Scrophulariaceae
.
Die natürlichen Pflanzenfamilien
4, 3b
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