Erect, annual or perennial herbs, or subshrubs, variously hairy
Leaves
with lower ones usually opposite or subopposite, upper ones usually alternate, sessile
Flowers
pedicellate, in a bracteate, one-sided cyme or in a branching system of ebracteate, very lax cymes
Calyx
deeply divided into 5 or 6 lobes; lobes ovate to lanceolate, with rounded to cordate bases, apices narrow to cuspidate, often winged; strongly accrescent in fruit
Corolla
blue or white; tube campanulate to funnel-shaped, throat naked; lobes 5 or 6, ovate-acuminate or broadly triangular, often spreading or reflexed
Stamens
arising on corolla tube, sessile or with very short, flattened filaments; anthers oblong or linear-lanceolate, with connectives prolonged above, the whole androecium converging above to form a cone with connective prolongations twisted together at apex, connectives with long, shaggy hairs on their dorsal surface
Ovary
4-lobed, 4-locular; gynobase pyramidal; style gynobasic or terminal, long and subulate; stigma small, subglobose
Fruit
of 4 ovoid nutlets, smooth or variously ornamented, or of a single, sessile nutlet; nutlets enclosed in enlarged calyx
x = 12, 11
Nomenclature:
Trichodesma
R.Br.
Brown: 496 (1810) name conserved
Lehmann: 192 (1818)
Candolle: 171 (1846)
Bentham: 845 (1876)
Boissier: 280 (1879)
Gürke: 99 (1897)
Baker & Wright: 44 (1906)
Brand: 19 (1921)
Riedl: 219 (1967)
Brummitt: 94 (1990)
Nowicke & Miller: 24 (1991)
Verdcourt: 91 (1991)
Borraginoides
Boehm.
Boehmer: 18 (1760).
Pollichia
Medik.
Medikus: 247 (1783).
Friedrichsthalia
Fenzl
Fenzl: 53 (1839).
Leiocarya
Hochst.
Hochstetter: 30 (1844)
Distribution & Notes:
Global
: Species ± 45, mainly in tropical and subtropical regions of Africa to Asia and Australia
Southern Africa
: Species 4, not recorded from Western Cape
References:
BAKER, J.G. & WRIGHT, C.H. 1906.
Boragineae
.
Flora of tropical Africa
4,2
BENTHAM, G. 1876.
Boragineae
. In G. Bentham & J.D. Hooker,
Genera plantarum
2. Reeve, London
BOEHMER, G.R 1760. Borraginioides. In C.G. Ludwig,
Definitiones generum plantarum
. Gleditsch, Leipzig
BOISSIER, P.E. 1879.
Borragineae
.
Flora orientalis
4. H. Georg, Geneva
BRAND, A. 1921.
Borraginaceae
-
Borraginoideae
/
Cynoglosseae
.
Das Pflanzenreich
78
BROWN, R. 1810.
Prodromus florae Novae Hollandiae
. J. Johnson & Co., London
CANDOLLE, A.P. DE. 1846.
Borragineae
.
Prodromus
10. Victor Masson, Paris
FENZL, E. 1839.
Friedrichsthalia
. In S.L. Endlicher & E. Fenzl,
Novarum stirpium decades
. Sollinger, Vienna
GÜRKE, M. 1897.
Borraginaceae
.
Die natürlichen Pflanzenfamilien
4, 3a
HOCHSTETTER, C.F. 1844. Nova genera plantarum Africae, proponit et describit.
Flora
27
LEHMANN, J.G.C. 1818.
Plantae e familia Asperifoliarum nuciferae
. Dümmler, Berlin
MEDIKUS, F.K. 1783.
Botanische Beobachtungen des Jahres 1783
. Hof- und akademische Buchhandlung, Mannheim
NOWICKE, J.W. & MILLER, J.S. 1991.
Boraginaceae
. In M.D. Dassanayake,
A revised handbook to the flora of Ceylon
. Amerind Publishing Co. Pvt. Ltd., New Delhi
RIEDL, H. 1967.
Boraginaceae
.
Flora iranica
48
VERDCOURT, B. 1991.
Boraginaceae
.
Flora of tropical East Africa
.
Boraginaceae
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