Trees, shrubs, or somewhat scandent, with or without axillary spines; dioecious, rarely monoecious or flowers bisexual
Leaves
mostly opposite, sometimes alternate or whorled, entire, petiolate
Inflorescences
paniculate cymes, axillary or terminal, dense or lax, subsessile or pedunculate
Flowers
unisexual (occasionally bisexual), small; bracts 2-4, not involucrate
Male flower
: perianth infundibuliform, campanulate, urceolate or tubular, 5-10-lobed or -toothed; tube with viscid glands on ribs; lobes or teeth induplicate-valvate, erect, spreading or reflexed; stamens usually 5-10, mostly exserted; filaments connate below into a tube or ring
Female flower
: perianth similar to male, but tube usually enlarged at base; stamens rudimentary; ovary sessile or stipitate, elongate; style filiform, included or exserted; stigma capitellate or peltate, pedicellate, multifid
Fruit
enclosed in hardened, elongated, oblong or clavate, smooth or 4- or 5-angled perianth base (anthocarp); anthocarp coriaceous, with longitudinal rows of viscid stipitate glands
Seed
with straight embryo
x = 8 (high polyploidy)
Nomenclature:
Pisonia
L.
Linnaeus: 1026 (1753)
Linnaeus: 451 (1754)
Jussieu: 91 (1789)
Brown: 422 (1810)
Choisy: 440 (1849)
Hooker: 9 (1880)
Heimerl: 29 (1889)
Baker & Wright: 8 (1909)
Cooke: 397 (1910)
Heimerl: 126 (1934)
Stemmerik: 457 (1964a)
Stemmerik: 275 (1964b)
Meikle & Hewson: 16 (1984)
Stannard: 25 (1988)
Bittrich & Kühn: 482 (1993)
Distribution & Notes:
Global
: Species ± 50, cosmopolitan in tropical and subtropical regions, predominantly of America and SE Asia, with a few species in Africa and Australia
Southern Africa
: Species 1:
Pisonia aculeata
L., N KwaZulu-Natal
References:
BAKER, J.G. & WRIGHT, C.H. 1909.
Nyctagineae
.
Flora of tropical Africa
6,1
BITTRICH, V. & KÜHN, U. 1993.
Nyctaginaceae
. In K. Kubitzki, J.G. Rohwer & V. Bittrich,
The families and genera of vascular plants - dicotyledons
2. Springer-Verlag, Berlin
BROWN, R. 1810.
Nyctagines
Juss.
Prodromus florae Novae Hollandiae et Insulae van-Diemen
1. Johnson & Co., London
CHOISY, J.D. 1849.
Nyctagineae
. In A. de Candolle,
Prodromus
13,2. V. Masson, Paris
COOKE, T. 1910.
Nyctagineae
.
Flora capensis
5,1
HEIMERL, A. 1889.
Nyctaginaceae
.
Die natürlichen Pflanzenfamilien
3,1b
HEIMERL, A. 1934.
Nyctaginaceae
.
Die natürlichen Pflanzenfamilien
, edn 2, 16c
HOOKER, J.D. 1880.
Nyctaginaceae
. In G. Bentham & J.D. Hooker,
Genera plantarum
3,1. Lovell Reeve & Co., London
JUSSIEU, A.L. DE. 1789. Nyctagines, les Nictages.
Genera plantarum secundum ordines naturales disposita
. Herissant & Barrois, Paris
LINNAEUS, C. 1753.
Species plantarum
, edn 1. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754.
Genera plantarum
, edn 5. Laurentius Salvius, Stockholm
MEIKLE, R.D. & HEWSON, H.J. 1984.
Nyctaginaceae
.
Flora of Australia
4
STEMMERIK, J.F. 1964b. Florae Malesianae Precursores 38. Notes on
Pisonia
L. in the Old World (
Nyctaginaceae
).
Blumea
12
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