e-Key v3 - Heliotropium
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Interactive keys to the identification of seed plants of southern Africa using keys based on plant morphology.

Boraginaceae - Heliotropioideae - Heliotropium L.

Description :

  • Annual or perennial herbs, sometimes subshrubs, rarely shrubs, usually hairy
  • Leaves alternate, rarely subopposite, sessile or petiolate, entire or denticulate
  • Flowers usually small, many, sessile, in unilateral, terminal or pseudoaxillary helicoid cymes with cymes simple or peduncle dichotomously branched to give a pair of cymes or flowers solitary at apices of leafy branches; bracts present or 0
  • Calyx with a short tube; 5-lobed, lobes lanceolate or linear
  • Corolla white, cream, yellow or orange yellow; tube usually cylindric, glabrous or hairy, throat often pubescent; lobes 5, variably shorter than tube, spreading
  • Disc present or 0
  • Stamens arising on corolla tube, included; filaments very short
  • Ovary completely or incompletely 4- or 5-locular with a single, pendulous ovule in each locule; gynobase flat; anthers elliptic-oblong or lanceolate, sometimes pubescent at apices; style terminal, undivided, included; stigma conical at apex, with a broad ring at base of cone
  • Fruit dry or somewhat fleshy, usually with 4 distinct 1-seeded nutlets, or nutlets cohering in pairs or reduced to 1 nutlet by abortion; nutlets with outer surface variable
  • x = 13, 8 (7) (aneuploids, high polyploidy)

Nomenclature:

  • Heliotropium L.
    • Linnaeus: 140 (1753)
    • Linnaeus: 63 (1754)
    • Candolle: 532 (1845)
    • Bentham: 843 (1876)
    • Boissier: 125 (1879)
    • Gürke: 90 (1897)
    • Wright: 7 (1904)
    • Friedrich-Holzhammer: 1 (1967)
    • Taton: 28 (1971)
    • Martins: 83 (1990)
    • Al-Shehbaz: 74 (1991)
    • Nowicke & Miller: 29 (1991)
    • Verdcourt: 50 (1991)
    • Riedl: 99 (1997)

Distribution & Notes:

  • Global : Species ± 250, in tropical to warm, temperate regions, particularly in ± arid areas
  • Southern Africa : Species 20, widespread but not recorded from Lesotho and Western Cape

References:

  • AL-SHEHBAZ, I.A. 1991. The genera of Boraginaceae in the southeastern United States. Journal of the Arnold Arboretum , supplementary series 1
  • BENTHAM, G. 1876. Boragineae . In G. Bentham & J.D. Hooker, Genera plantarum 2. Reeve, London
  • BOISSIER, P.E. 1879. Borragineae . Flora orientalis 4. H. Georg, Geneva
  • CANDOLLE, A.P. DE. 1845. Borragineae . Prodromus 9. Treuttel & Würtz, Paris
  • FRIEDRICH-HOLZHAMMER, M. 1967. Heliotropiaceae . Prodromus einer Flora von Südwestafrika 119
  • GÜRKE, M. 1897. Borraginaceae . Die natürlichen Pflanzenfamilien 4, 3a
  • LINNAEUS, C. VON. 1753. Species plantarum . Laurentius Salvius, Stockholm
  • LINNAEUS, C. VON. 1754. Genera plantarum , edn 5. Laurentius Salvius, Stockholm
  • MARTINS, E.S. 1990. Boraginaceae . Flora zambesiaca 7,4
  • NOWICKE, J.W. & MILLER, J.S. 1991. Boraginaceae . In M.D. Dassanayake, A revised handbook to the flora of Ceylon . Amerind Publishing Co. Pvt. Ltd., New Delhi
  • RIEDL, H. 1997. Boraginaceae . Flora malesiana 13
  • TATON, A. 1971. Boraginaceae . Flore du Congo du Rwanda et du Burundi . Boraginaceae
  • VERDCOURT, B. 1991. Boraginaceae . Flora of tropical East Africa . Boraginaceae
  • WRIGHT, C.H. 1904. Boragineae . Flora capensis 4,2