Annual or perennial herbs, sometimes subshrubs, rarely shrubs, usually hairy
Leaves
alternate, rarely subopposite, sessile or petiolate, entire or denticulate
Flowers
usually small, many, sessile, in unilateral, terminal or pseudoaxillary helicoid cymes with cymes simple or peduncle dichotomously branched to give a pair of cymes or flowers solitary at apices of leafy branches; bracts present or 0
Calyx
with a short tube; 5-lobed, lobes lanceolate or linear
Corolla
white, cream, yellow or orange yellow; tube usually cylindric, glabrous or hairy, throat often pubescent; lobes 5, variably shorter than tube, spreading
Disc
present or 0
Stamens
arising on corolla tube, included; filaments very short
Ovary
completely or incompletely 4- or 5-locular with a single, pendulous ovule in each locule; gynobase flat; anthers elliptic-oblong or lanceolate, sometimes pubescent at apices; style terminal, undivided, included; stigma conical at apex, with a broad ring at base of cone
Fruit
dry or somewhat fleshy, usually with 4 distinct 1-seeded nutlets, or nutlets cohering in pairs or reduced to 1 nutlet by abortion; nutlets with outer surface variable
x = 13, 8 (7) (aneuploids, high polyploidy)
Nomenclature:
Heliotropium
L.
Linnaeus: 140 (1753)
Linnaeus: 63 (1754)
Candolle: 532 (1845)
Bentham: 843 (1876)
Boissier: 125 (1879)
Gürke: 90 (1897)
Wright: 7 (1904)
Friedrich-Holzhammer: 1 (1967)
Taton: 28 (1971)
Martins: 83 (1990)
Al-Shehbaz: 74 (1991)
Nowicke & Miller: 29 (1991)
Verdcourt: 50 (1991)
Riedl: 99 (1997)
Distribution & Notes:
Global
: Species ± 250, in tropical to warm, temperate regions, particularly in ± arid areas
Southern Africa
: Species 20, widespread but not recorded from Lesotho and Western Cape
References:
AL-SHEHBAZ, I.A. 1991. The genera of
Boraginaceae
in the southeastern United States.
Journal of the Arnold Arboretum
, supplementary series 1
BENTHAM, G. 1876.
Boragineae
. In G. Bentham & J.D. Hooker,
Genera plantarum
2. Reeve, London
BOISSIER, P.E. 1879.
Borragineae
.
Flora orientalis
4. H. Georg, Geneva
CANDOLLE, A.P. DE. 1845.
Borragineae
.
Prodromus
9. Treuttel & Würtz, Paris
FRIEDRICH-HOLZHAMMER, M. 1967.
Heliotropiaceae
.
Prodromus einer Flora von Südwestafrika
119
GÜRKE, M. 1897.
Borraginaceae
.
Die natürlichen Pflanzenfamilien
4, 3a
LINNAEUS, C. VON. 1753.
Species plantarum
. Laurentius Salvius, Stockholm
LINNAEUS, C. VON. 1754.
Genera plantarum
, edn 5. Laurentius Salvius, Stockholm
NOWICKE, J.W. & MILLER, J.S. 1991.
Boraginaceae
. In M.D. Dassanayake,
A revised handbook to the flora of Ceylon
. Amerind Publishing Co. Pvt. Ltd., New Delhi
RIEDL, H. 1997.
Boraginaceae
.
Flora malesiana
13
TATON, A. 1971.
Boraginaceae
.
Flore du Congo du Rwanda et du Burundi
.
Boraginaceae
VERDCOURT, B. 1991.
Boraginaceae
.
Flora of tropical East Africa
.
Boraginaceae
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