Annual or perennial herbs; stems erect, ascending, prostrate or decumbent, often rooting at nodes and much branched, sometimes fleshy and/or hollow, glabrous or pubescent
Leaves
usually opposite, especially near base, cauline ones sometimes alternate, verticillate or scattered, sessile or petiolate, ovate to lanceolate-oblong, entire or variously dentate or crenate
Flowers
in terminal or axillary racemes, rarely axillary and solitary, bracteate, pedicellate; bracts 0
Calyx
with very short tube, 4(5)-partite, where present, fifth posterior lobe smaller, accrescent
Corolla
barely tubular, usually subrotate, 4(5)-lobed; tube very short, rarely exceeding calyx; limb spreading, lateral lobes exterior in bud
Stamens
2, fertile, arising in corolla tube at sides of upper lobe, exserted; anthers bithecate; thecae divergent or parallel, confluent at apex
Ovary
bilocular; ovules few to many; style with subcapitate stigma
Fruit
a laterally compressed or turgid, bivalved, loculicidal (sometimes septicidal) capsule, ovoid to reniform or obcordate
Seeds
ovoid or orbicular
x = 7, 8, 9, 17 (6, 10, 12, 15) (B-chromosomes, aneuploids, high polyploidy)
Nomenclature:
Veronica
L.
Linnaeus: 9 (1753)
Hiern: 367 (1904)
Henderson & Anderson: 292 (1966)
Merxmüller & Roessler: 56 (1967)
Walters & Webb: 242 (1972)
Wells et al.: 536 (1986)
Philcox: 80 (1990)
Distribution & Notes:
Global
: Species ± 180, mostly temperate, mainly north, few on tropical mountains in Africa, some subcosmopolitan weeds
Southern Africa
: Species 1:
Veronica anagallis-aquatica
L., indigenous or introduced from Europe, widespread in wet areas but not Botswana; several others as sporadic escapes or weeds, especially *
V. persica
Desf
References:
HENDERSON, M. & ANDERSON, J.G. 1966. Common weeds in South Africa.
Memoirs of the Botanical Survey of South Africa
No. 37
LINNAEUS, C. 1753.
Species plantarum
, edn 1. Laurentius Salvius, Stockholm
MERXMÜLLER, H. & ROESSLER, H. 1967.
Scrophulariaceae
.
Prodromus einer Flora von Südwestafrika
126
PHILCOX, D. 1990.
Scrophulariaceae
.
Flora zambesiaca
8,2
WALTERS, S.M. & WEBB, D.A. 1972.
Veronica
L. In D.A. Webb,
Scrophulariaceae
.
Flora europaea
3
WELLS, M.J., BALSINHAS, A.A., JOFFE, H., ENGELBRECHT, V.M., HARDING, G. & STIRTON, C.H. 1986. A catalogue of problem plants in southern Africa.
Memoirs of the Botanical Survey of South Africa
No. 53
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