Evergreen trees, shrubs or rhizomatous geoxylic suffrutices; wood with abundant silica inclusions
Leaves
alternate, simple, entire, coriaceous, lower surface with close, prominent, reticulate venation which delimits small stomatal cavities filled with hairs, sometimes with 2 glands on upper side of petiole or at junction of petiole with blade; stipules subulate or lanceolate, caducous or persistent
Inflorescences
axillary, many-flowered, complex cymes or cymose panicles; bracts and bracteoles eglandular, completely concealing flower bud
Calyx
: tube (receptacle) turbinate and often obliquely bent or cup-shaped and ± unilaterally gibbous, hairy inside throughout; lobes 5, free, obtuse and imbricate or triangular-acute and sometimes subvalvate
Petals
5, free, inserted in mouth of perianth tube, sessile or clawed, caducous
Stamens
6-10 or more, sometimes unequal, inserted in throat of perianth tube, connate at base into a short ring or unilateral bundle, sometimes almost free, about as long as perianth lobes; filaments subulate-filiform, white; anthers small, dorsifixed, 2-thecous, dehiscing longitudinally and introrsely
Ovary
superior, adnate to one side of perianth tube; carpels 1, rarely 2 or 3, 2-locular, with a single, erect ovule in each locule, often villous; style arising at base of carpel(s), terete, often hairy, arcuate, included; stigma ± truncate
Fruit
drupaceous, 1-seeded, ellipsoid, obovoid or globose; epicarp verrucose; endocarp hard, thick, with a rough, fibrous surface, with 2 basal obturators
Seed
erect; endosperm 0
x = 11 (polyploidy)
Nomenclature:
Chrysobalanaceae
Brown: 433 (1818) as
Chrysobalaneae
Candolle: 525 (1825) as
Rosaceae
tribus
Chrysobalaneae
Endlicher: 1251 (1840) as
Rosaceae
order
Chrysobalaneae
Hooker: 601 (1865) as
Rosaceae
Focke: 55 (1894) as
Rosaceae
subfamily
Chrysobalanoideae
Prance: 1 (1972) as
Chrysobalanaceae
White: 265 (1976)
Prance & White: 1 (1988)
Distribution & Notes:
Global
: Genera 17, species ± 490; restricted to the lowlands of the tropical and subtropical regions of both hemispheres, concentrated near the equator, especially in the Neotropics
Southern Africa
: Genus 1, species 2
References:
BROWN, R. 1818.
Chrysobalaneae
. In J.H.Tuckey,
Narrative of an expedition to explore the river Zaire, usually called the Congo
. Appendix 5. John Murray, London
CANDOLLE, A.-P. DE. 1825.
Rosaceae
tribus
Chrysobalaneae
.
Prodromus
2. Treuttel & Würtz, Paris
ENDLICHER, S.L. 1840.
Rosaceae
order
Chrysobalaneae
.
Genera plantarum secundum ordines naturales disposita
2. Beck, Vienna
PRANCE, G.T. & WHITE, F. 1988. The genera of
Chrysobalanaceae
: a study in practical and theoretical taxonomy and its relevance to evolutionary biology.
Philosophical Transactions of the Royal Society, London
B 320
WHITE, F. 1976. The taxonomy, ecology and chorology of African
Chrysobalanaceae
(excluding
Acioa
).
Bulletin du Jardin Botanique National du Belgique
46
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