Euphorbiaceae
-
Phyllanthoideae
-
Phyllantheae
-
Phyllanthus
L.
Description
:
Herbs, undershrubs from perennial rootstocks, shrubs, or trees; monoecious or sometimes dioecious
Leaves
scale-like cataphylls or foliage leaves, alternate; stipulate
Inflorescence
mainly of many fascicled male flowers and solitary female flowers in upper axils
Petals
0
Male flowers
: sepals 4-6, membranous, free or slightly connate at base, imbricate; disc often small, of separate glands, rarely annular; stamens 2-6, in middle of flower; filaments free or connate, or some free and some connate; ovary 0
Female flowers
: sepals as in male or larger; disc usually annular, saucer- or cup-shaped, rarely of separate glands; ovary 3(-many)-locular, with 2 ovules in each locule; styles 3, free or partly connate, usually bifid or bilobed
Fruit
a capsule, dry or somewhat fleshy, dehiscent or subdehiscent; endocarp usually crustaceous
Seeds
3-angled, often longitudinally sulcate or pitted; testa thinly crustaceous; albumen fleshy; embryo straight or slightly incurved; cotyledons flat, straight or rarely flexuous
x = 13 (6, 7, 8, 9, 10, 11, 12) (aneuploids, high polyploidy, B-chromosomes)
Nomenclature:
Phyllanthus
L.
Linnaeus: 987 (1753)
Hutchinson: 386 (1920)
Radcliffe-Smith: 10 (1987)
Webster: 44 (1994)
Radcliffe-Smith: 36 (1996)
Distribution & Notes:
Global
: Species 750-800; cosmopolitan in tropical and subtropical rcgions, ± 150 in Africa
Southern Africa
: Species 22, widespread
References:
HUTCHINSON, J. 1920.
Euphorbiaceae
[in part].
Flora capensis
5, 2
LINNAEUS, C. 1753.
Species plantarum
, edn 1. Laurentius Salvius, Stockholm
RADCLIFFE-SMITH, A. 1987.
Euphorbiaceae
.
Flora of tropical East Africa
.
Euphorbiaceae
Part 1
RADCLIFFE-SMITH, A. 1996.
Euphorbiaceae
.
Flora zambesiaca
9, 4
WEBSTER, G.L. 1994. Synopsis of the genera and suprageneric taxa of
Euphorbiaceae
.
Annals of the Missouri Botanical Garden
81
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