Annual, biennial or perennial herbs, sometimes woody at base, diffuse, often glandular-viscid; stems often decumbent or ascending, swollen at nodes, with forked or fascicled branches
Leaves
opposite, sometimes ± fleshy, linear to filiform, with abbreviated leaf buds in their axils and appearing whorled; stipules scarious, connate in pairs and sheathing the node
Inflorescences
cymose or pseudo-racemose
Flowers
bisexual, rarely unisexual, subperigynous
Sepals
5, free, divided almost to base, obscurely nerved, with scarious margins
Petals
5, entire, rarely 0, white, pink or red
Stamens
(2)5-10, unequal, arising on a perigynous ring; anthers medifixed, introrse
Ovary
1-locular; ovules many on free-central placenta; styles 3(-5), free or ± connate
Fruit
a capsule, subglobose or ovoid-trigonous, dehiscent almost to base into 3(-5) valves or teeth
Seeds
many, ovoid, pyriform or subtrigonous, often winged; testa smooth, rugose or variously tuberculate; embryo annular or hooked
x = 9 (aneuploids, high polyploidy)
Nomenclature:
Spergularia
(Pers.) J.Presl & C.Presl
Presl & Presl: 94 (1819)
Pax & Hoffmann: 312 (1934)
Monnier & Ratter: 154 (1964)
Adamson: 394 (1950)
Meikle: 274 (1977)
Friedrich: 782 (1979)
Bittrich: 220 (1993)
Arenaria
section
Spergularia
Pers.
Persoon: 504 (1805).
Lepigonum
Sond.
Sonder: 134 (1860)
Distribution & Notes:
Global
: Species ± 25, cosmopolitan, especially some halophytes with nearly worldwide distribution
Southern Africa
: Species 4, mainly in Cape region, but also scattered in Namibia, North-West, Free State and KwaZulu-Natal
References:
ADAMSON, R.S. 1950.
Caryophyllaceae
. In R.S. Adamson & T.M. Salter,
Flora of the Cape Peninsula
. Juta, Cape Town
BITTRICH, V. 1993.
Caryophyllaceae
. In K. Kubitzki, J.G. Rohwer & V. Bittrich,
The families and genera of vascular plants - dicotyledons
2. Springer-Verlag, Berlin
FRIEDRICH, H.C. 1979.
Caryophyllaceae
. In G. Hegi,
Illustrierte Flora von Mitteleuropa
, edn 2; 3,2
MEIKLE, R.D. 1977.
Caryophyllaceae
.
Flora of Cyprus
1
MONNIER, P. & RATTER, J.A. 1964.
Caryophyllaceae
. Subfamily
Paronychioideae
.
Spergularia
.
Flora europaea
1
PAX, F. & HOFFMANN, K. 1934.
Caryophyllaceae
.
Die natürlichen Pflanzenfamilien
II,16c
PERSOON, C.H. 1805.
Synopsis plantarum
1. Treuttel & Würtz, Paris
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