Erect, annual or short-lived perennial herbs, with long stinging hairs on all aerial parts; plants monoecious or dioecious by abortion
Leaves alternate, elliptic to ovate, mostly variously divided, coarsely serrate or lobed, always 3-nerved, petiolate, with dot-like cystoliths; stipules intrapetiolar, connate, often large
Inflorescences simple or paniculate spikes, axillary and solitary or paired, male often long, slender with flowers clustered on rachis; female sessile, very densely branched, armed with stinging hairs
Male flowers pedicellate; tepals 4 or 5, ovate, concave, valvate in bud; bud depressed; stamens as many as tepals; ovary rudimentary
Female flowers sessile, with 3 almost completely connate tepals ± enclosing ovary, split to base on one side, sometimes with 1 minute free tepal; staminodes 0; ovary ± asymmetrical, reflexed, laterally compressed; ovule erect from base; stigma subulate to filiform, acute, very minutely papillose
Achene broad, oblique, much compressed, rugose, with remains of persistent perianth at base
x = 10, 12 (B-chromosomes)
Nomenclature:
Girardinia Gaudich.
Gaudichaud-Beaupré: 498 (1830)
Endlicher: 283 (1837)
Weddell: 163, t. 2B (1856)
Weddell: 100 (1869)
Bentham: 384 (1880)
Engler: 107 (1888)
Rendle: 265 (1917)
Friis: 143 (1981)
Friis: 12 (1989)
Friis: 85 (1991)
Friis: 621 (1993)
Distribution & Notes:
Global: Species 2, mountains of Old World tropics, from West Africa, Madagascar, Asia to SE China
Southern Africa: Species 1: Girardinia diversifolia (Link) Friis, Northern Province and Mpumalanga
References:
BENTHAM, G. 1880. Tribus Urticeae. In G. Bentham & J.D. Hooker, Genera plantarum 3. Lovell Reeve & Co., London
ENDLICHER, S.L. 1837. Urticaceae. Genera plantarum secundum ordines naturales disposita. Beck, Vienna
ENGLER, A. 1888. Urticaceae. Die natürlichen Pflanzenfamilien 3,1
FRIIS, I. 1981. A synopsis of Girardinia (Urticaceae). Kew Bulletin 36
FRIIS, I. 1989. Flora of tropical East Africa. Urticaceae
FRIIS, I. 1991. Urticaceae. Flora zambesiaca 9,6
FRIIS, I. 1993. Urticaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
GAUDICHAUD-BEAUPRÉ, C. 1830. In M.L. de Freycinet, Voyage autour du monde ... exécuté sur les corvettes de S.M. l'Uranie et la Physicienne. Botanique. Pillet-ainé, Paris
RENDLE, A.B. 1917. Urticaceae. Flora of tropical Africa 6,2
WEDDELL, H.A. 1856. Monographie de la famille des Urticacées. Gide & J. Baudry, Paris. Reprinted in Archives du Muséum d'Histoire Naturelle, Paris 9
WEDDELL, H.A. 1869. Urticaceae. In A. de Candolle, Prodromus 16,1. Treuttel & Würtz, Paris
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