Adventitious roots developing from leaf scars of submerged leaves containing chlorophyll, pinnatisect into many filiform segments
Leaves dimorphic; floating leaves alternate, forming rosettes, only present at upper nodes, with long spongy petiole ± inflated about middle, lamina deltoid to rhombic, upper margins toothed; submerged leaves opposite, sessile, linear, entire, falling off early; stipules small, scarious, divided to base and thus apparently more than 2 per leaf, linear, elongate, acuminate, deciduous
Flowers bisexual, regular, solitary, axillary
Calyx 4-lobed; tube oblong, slightly produced above ovary; lobes accrescent, often spinescent, hirsute at first, glabrescent
Petals 4, free, subsessile, obovate, white
Stamens 4, as long as calyx lobes; anthers oblong, 2-thecous
Ovary half-inferior, 2-locular, with a single, pendulous, linear ovule in each locule; style thick, terete, as long as stamens; stigma globose
Fruit a 1-seeded, top-shaped, bony or coriaceous drupe, horned with 2-4 persistent calyx lobes
Seed solitary, filling fruit; testa woody; cotyledons very unequal, 1 almost vestigial
x = 6 or 12 (polyploidy)
Nomenclature:
Trapaceae
Linnaeus: 120 (1753)
Roessler 96 (1966)
Brenan: 346 (1978)
Verdcourt: 447 (1986)
Verdcourt: 11 (1998)
Distribution & Notes:
Global: Genus 1, Eurasia and Africa; 1 polymorphic species or ± 15, depending on species interpretation
Southern Africa: Genus 1, species 1 with 2 varieties
References:
BRENAN, J.P.M. 1978. Trapa L. Flora zambesiaca 4
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
ROESSLER, H. 1966. Trapaceae. Prodromus einer Flora von Südwestafrika 96
VERDCOURT, B. 1986. Notes on Trapa for the 'Flora of southern Africa.' Kew Bulletin 41
VERDCOURT, B. 1998. FSA contributions 10: Trapaceae. Bothalia 28
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