Annual, biennial or perennial herbs, sometimes with thickened, ± woody, short, basal stems, occasionally soft-wooded shrubs or undershrubs, rarely small trees, often with most parts rather succulent; rarely dioecious
Leaves opposite or alternate, simple, usually with entire margins, glabrous or with glandular, barbulate or simple hairs; stipules 0 or leaf axils with hairs, spines, bristles or scales, often interpreted as stipules
Inflorescences variously cymose and terminal, rarely raceme-like paniculate, or terminal heads of sessile flowers surrounded by an involucre of leaves, occasionally reduced to solitary axillary flowers
Flowers bisexual or occasionally unisexual, usually regular
Sepals 2(3), usually unequal, imbricate, deciduous or rarely persistent, free or occasionally adnate to base of ovary
Petals (2-4)5(-15), imbricate, free or sometimes connate near base, ephemeral, fugacious or persistent, sometimes accrescent
Stamens (3)4-many, often as many as petals, hypogynous, perigynous or sometimes epigynous, free from each other but sometimes adnate to petals at base, or sometimes epipetalous; anthers basifixed, opening through longitudinal slits, sometimes staminodal
Ovary superior, rarely half-inferior, with 2-5 carpels but 1 locule and basal or free-central placentation at maturity; ovules 1-many, often on long funicles; stigmas 2-5(-8), free or connate into a style
Fruit a capsule dehiscing through 3-5(-8) terminal valves or circumscissile, sometimes with epicarp and endocarp separating and sometimes with epicarp deciduous; sometimes a nut which may dehisce irregularly near base
Seeds sometimes arillate and aril sometimes enveloping seed, sometimes compressed and rarely winged, shiny or granulated, with or without appendages; testa usually hard and crustaceous; endosperm rudimentary; perisperm usually present and often abundant; embryo ± curved almost into a circle
Nomenclature:
Portulacaceae
Jussieu: 312 (1789)
Candolle: 354 (1828)
Fenzl: 946 (1839)
Bentham: 155 (1862)
Pax: 51 (1889)
Franz: 1 (1908)
Pax & Hoffmann: 234 (1934)
MacNeill: 725 (1974)
Nyananyo: 237 (1990)
Carolin: 544 (1993)
Gilbert: 120 (1993)
Distribution & Notes:
Global: Genera ± 30, species ± 450, largely a southern hemisphere family (Africa, Madagascar, Asia, Australia, New Zealand) extending into North America but poorly represented in Eurasia; some species widespread weeds
Southern Africa: Genera 7 (1 exotic), species 46 (3 exotic)
References:
BENTHAM, G. 1862. Portulaceae. In G. Bentham & J.D. Hooker, Genera plantarum 1. Lovell Reeve & Co., London
CANDOLLE, A.P. DE. 1828. Portulaceae. Prodromus 3. Treuttel & Würtz, Paris
CAROLIN, R.C. 1993. Portulacaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
FENZL, E. 1839. Ordo Portulacaceae. In S.L. Endlicher, Genera plantarum secundum ordines naturales disposita 2. Beck, Vienna
FRANZ, E. 1908. Beiträge zur Kenntnis der Portulacaceen und Basellaceen. Botanische Jahrbücher 42, Beiblatt 97
GILBERT, M.G. 1993. Portulacaceae. Flora of Somalia 1
JUSSIEU, A.L. DE. 1789. Portulaceae, les Portulacées. Genera plantarum secundum ordines naturales disposita. Herissant & Barrois, Paris
MCNEILL, J. 1974. Synopsis of a revised classification of the Portulacaceae. Taxon 23
NYANANYO, B.L. 1990. Tribal and generic relationship in the Portulacaceae (Centrospermae). Feddes Repertorium 101
PAX, F. 1889. Portulacaceae. Die natürlichen Pflanzenfamilien 3,1b
PAX, F. & HOFFMANN, K. 1934. Portulacaceae. Die natürlichen Pflanzenfamilien, edn 2,16c
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Biodiversity Advisor, developed by the South African National Biodiversity Institute (SANBI) and its Data Partners, is a system that will provide integrated biodiversity information to a wide range of users who will have access to geospatial data, plant and animal species distribution data, ecosystem-level data, literature, images and metadata.
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