Caryophyllaceae - Caryophylloideae - Caryophylleae - Dianthus L.
Description:
Annual or short-lived perennial herbs or subshrubs, tufted, often somewhat woody towards base; stems mostly thickened at nodes
Leaves opposite, grass-like, connate basally and forming a sheath, frequently 5-nerved; stipules 0
Inflorescences terminal panicles of cymes, in heads or occasionally flowers solitary, often subtended by bracts
Flowers bisexual, rarely unisexual; anthophore often present
Calyx gamosepalous, surrounded at base by 2-many bracts, tubular, firm, finely striate, 5-toothed, with many parallel veins, without membranous commissures
Petals 5, white, pink or red, rarely yellow, limb entire, toothed or fimbriate, without coronal scales; claw long, with two longitudinal ridges
Stamens 10; filaments long, slender
Ovary 1-locular, on a short gynophore, with chartaceous cap; ovules many on free central placenta; styles 2, linear, longer than ovary
Fruit a capsule, ovoid to cylindric, opening by 4 valves
Seeds many, orbicular to ovoid in outline, ± compressed, with facial hilum, black; testa with raised polygonal sculpturing; embryo straight
x = 15 (high polyploidy)
Nomenclature:
Dianthus L.
Linnaeus: 409 (1753)
Linnaeus: 191 (1754)
Jussieu: 302 (1789)
Candolle: 355 (1824)
Sonder: 122 (1860)
Pax: 76 (1889)
Williams: 346 (1893)
Burtt Davy: 148 (1926)
Pax & Hoffmann: 356 (1934)
Hooper: 1 (1959)
Tutin: 188 (1964)
Podlech: 1 (1967)
Wild, Hooper & Diniz: 16 (1973)
Meikle: 214 (1977)
Bittrich: 232 (1993)
Distribution & Notes:
Global: Species ± 300, Europe, Asia, especially Mediterranean, Africa
Southern Africa: Species 15, widely distributed. Some species are cultivated as ornamentals (e.g. *Dianthus caryophyllus L. and * D. chinensis L.)
References:
BITTRICH, V. 1993. Caryophyllaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
BURTT DAVY, J. 1926. Caryophyllaceae. A manual of the flowering plants and ferns of the Transvaal with Swaziland 1. Longmans, London
CANDOLLE, A.-P. DE. 1824. Caryophylleae. Prodromus 1. Treuttel & Würtz, Paris
HOOPER, S.S. 1959. The genus Dianthus in Central and South Africa. Hooker's Icones Plantarum, fifth series, 7,1
JUSSIEU, A.L. DE. 1789. Caryophylleae, les caryophyllées. Genera plantarum secundum ordines naturales disposita. Herissant & Barrois, Paris
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754. Genera plantarum, edn 5. Laurentius Salvius, Stockholm
MEIKLE, R.D. 1977. Caryophyllaceae. Flora of Cyprus 1
PAX, F. 1889. Caryophyllaceae. Die natürlichen Pflanzenfamilien 3,1b
PAX, F. & HOFFMANN, K. 1934. Caryophyllaceae. Die natürlichen Pflanzenfamilien II,16c
PODLECH, D. 1967. Caryophyllaceae. Prodromus einer Flora von Südwestafrika 31
WILD, H., HOOPER, S.S. & DINIZ, M.A. 1973. Caryophyllaceae. Flora de Moçambique 19
WILLIAMS, F.N. 1893. A monograph of the genus Dianthus L. The Journal of the Linnean Society (Botany) 29
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