Amaranthaceae - Amaranthoideae - Amarantheae - Aervinae - Achyranthes L.
Description:
Erect perennial herbs, sometimes bushy or scandent, sometimes flowering in first year
Leaves opposite, entire, petiolate
Inflorescence a ± slender, simple or branched, elongated solitary or fasciculate spike up to 0.3 m long, terminal on stem and branches, flowers at first congested and ± patent, finally usually laxer and deflexed; bracts deltoid, ovate or acuminate, midrib excurrent in a spine, deflexed in fruit, persistent
Flowers bisexual, solitary in axils of bracts; bracteoles 2, spinous-aristate with excurrent midrib, lamina forming short and free to longer and adnate membranous wings; entire flower with bracteoles falling with ripening of seed
Tepals 4 or 5, 1-3(-5)-nerved, narrowly lanceolate, acuminate, mucronate with excurrent midrib, indurated in fruit especially at base
Stamens 2-5; filaments filiform, monadelphous at base, alternating with usually fringed pseudostaminodes; anthers 2-thecous
Ovary turbinate or obovoid; ovule solitary from apex of an elongated basal funicle; style cylindrical, variable in length; stigma capitellate
Capsule thin-walled, indehiscent
Seed cylindrical; perisperm/endosperm copious
x = 7 (aneuploids, high polyploidy)
Nomenclature:
Achyranthes L.
Linnaeus: 204 (1753)
Linnaeus: 96 (1754)
Brown: 417 (1810)
Hooker: 35 (1880)
Schinz: 112 (1893)
Baker & Clarke: 62 (1909)
Cooke & Wright: 427 (1910)
Adamson: 361 (1950)
Bullock: 73 (1957)
Cavaco: 302 (1958)
Cavaco: 114 (1962)
Podlech: 4 (1966)
Henderson & Anderson: 102 (1966)
Townsend: 145 (1973)
Townsend: 473 (1974)
Townsend: 100 (1985)
Townsend: 105 (1988)
Townsend: 163 (1993a)
Townsend: 87 (1993b)
Distribution & Notes:
Global: Species 6-8, warm temperate and tropical regions of the Old World
Southern Africa: Species 1: Achyranthes aspera L. with 3 varieties, widespread and very variable in robustness
References:
ADAMSON, R.S. 1950. Amaranthaceae Lindl. In R.S. Adamson & T.M. Salter, Flora of the Cape Peninsula. Juta, Cape Town
BAKER, J.G. & CLARKE, C.B. 1909. Amarantaceae. Flora of tropical Africa 6,1
BROWN, R. 1810. Amarantaceae Juss. Prodromus florae novae Hollandiae et Insulae van-Diemen 1. Johnson & Co., London
BULLOCK, A.A. 1957. The application of the generic name Achyranthes. Kew Bulletin 1957
CAVACO, A. 1958. Les Achyranthes (Amaranthaceae) du continent Africain et des îles adjacentes. Boletim da Sociedade Broteriana 32
CAVACO, A. 1962. Les Amaranthaceae de l'Afrique au sud du Tropique du Cancer et de Madagascar. Mémoires du Muséum National d'Histoire Naturelle, Paris, Série B, Botanique 13
COOKE, T. & WRIGHT, C.H. 1910. Amarantaceae. Flora capensis 5,1
HENDERSON, M.D. & ANDERSON, J.G. 1966. Common weeds in South Africa. Memoirs of the Botanical Survey of South Africa 37
HOOKER, J.D. 1880. Amarantaceae. In G. Bentham & J.D. Hooker, Genera plantarum 3,1. Lovell Reeve & Co., London
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754. Genera plantarum, edn 5. Laurentius Salvius, Stockholm
PODLECH, D. 1966. Amaranthaceae. Prodromus einer Flora von Südwestafrika 33
SCHINZ, H. 1893. Amarantaceae. Die natürlichen Pflanzenfamilien III,1a
TOWNSEND, C.C. 1973. Notes on Amaranthaceae 1. Kew Bulletin 28
TOWNSEND, C.C. 1974. Notes on Amaranthaceae 2. Kew Bulletin 29
TOWNSEND, C.C. 1985. Flora of tropical East Africa. Amaranthaceae
TOWNSEND, C.C. 1993a. Amaranthaceae. Flora of Somalia 1
TOWNSEND, C.C. 1993b. Amaranthaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
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