Annual or perennial herbs or shrublets, generally erect, sometimes prostrate, usually woolly or cobwebby, often glandular, rarely glabrous
Leaves alternate, straight, variously shaped, sessile or petiolate, often hairy, pilose to velutinous, margins entire, flat or sometimes revolute
Capitula disciform or discoid; cylindric, turbinate, campanulate, subglobose or depressed-globose, in terminal, loose, flat-topped corymbs or solitary
Involucral bracts in few to many rows; stereome usually fenestrated (divided); white, yellow, straw-coloured, brown, pink, purple, or red, sometimes a combination of colours; glabrous or hairy
Receptacle smooth, honeycombed or fimbrilliferous, flat, epaleate, rarely paleate
Marginal florets female, fewer or sometimes more than central florets; corolla often narrowly tubular, sometimes with well-developed lamina, rarely filiform, yellow
Central florets bisexual; corolla funnel-shaped or tubular and then ± campanulate above, lobes 5, hairs always present on backs of lobes, yellow
Anthers ecalcarate, caudate; with lanceolate apical appendages; endothecial tissue polarised
Style branches truncate, penicillate; stigmatic areas separated
Cypselas sparsely hairy with short, clavate, myxogenic hairs or with long nonmyxogenic hairs or glabrous; epidermis smooth or often papillose
Pappus of barbellate or subplumose bristles, generally in 1 row; bristles basally with or without patent cilia, apical cells clavate or not
x = 7 (4, 5, 6, 8, 13) (aneuploids, high polyploidy)
Nomenclature:
Helichrysum Mill.
Miller: [462] (1754)
Persoon: 414 (1807) name conserved
Hilliard: 61 (1983)
Anderberg: 141 (1991)
Anderberg: 358 (1994)
Achyrocline (Less.) DC.
Hilliard & Burtt: 201 (1981)
Hilliard: 59 (1983)
Mesfin Tadesse & Reilly: 379 (1995).
Leontonyx Cass.
Cassini: 466 (1822)
Distribution & Notes:
Global: Species ± 600, largely Africa and Madagascar, also Europe, Asia and Australia
Southern Africa: Species 244, widely distributed
References:
ANDERBERG, A.A. 1991. Taxonomy and phylogeny of the tribe Gnaphalieae (Asteraceae). Opera Botanica 104
ANDERBERG, A.A. 1994. Tribes Inuleae, Plucheeae, Gnaphalieae. In K. Bremer, Asteraceae, cladistics and classification. Timber Press, Portland, Oregon
CASSINI, A.H.G. DE. 1822. Dictionnaire des sciences naturelles 25. Ed. F. Cuvier. edn 2. Le Normant, Paris
HILLIARD, O.M. 1983. Asteraceae (Compositae), Tribe Inuleae subtribe Gnaphaliinae (first part). Flora of southern Africa 33, 7, 2: 59, 61
HILLIARD, O.M. & BURTT, B.L. 1981. Some generic concepts in Compositae - Gnaphaliinae. Botanical Journal of the Linnean Society 82
MESFIN TADESSE & REILLY, T. 1995. 17. A contribution to studies on Helichrysum (Compositae - Gnaphalieae) - a revision of the species of north-east tropical Africa. In D.J.H. Hind, C. Jeffrey & G.V. Pope, Advances in Compositae Systematics. Royal Botanic Gardens, Kew
MILLER, P. 1754. The gardener's dictionary (Abridged edition), edn 4. Rivington, London
PERSOON, C.H. 1807. Helichrysum. Synopsis plantarum. 2. Cramer, Paris
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