Compact, ± stemless, dwarf, succulent perennials, ± sunken in the soil, of 1 or more ± connate leaf pairs (bodies); roots fibrous
Leaves opposite, connate to form an obconic body with a truncate apex, old leaves turning into ± hard sheaths enclosing new body, flat tops variously marked, sculptured or windowed; surface ± smooth, but epidermal cells often bluntly elevated, rarely papillate, with remarkable, thick outer epidermal walls encrusted with cutinised blocks and thick crystal layers, crystals increasing basipetally in size
Flowers solitary, rarely 2 or 3, from centre of fissure between leaves of a pair, shortly pedicellate, ebracteate, 15-40 mm in diameter; opening mid-afternoon, closing at sunset
Sepals 4-7
Petals 1-4-seriate, linear, white, yellow or orange
Stamens sometimes papillate towards base; staminodes 0
Nectary a crenulate ring
Ovary with parietal placentas; style short or 0; stigmas 4-7(9), filiform
Fruit a 4-7(9)-locular capsule, of Delosperma type; small; valves with broad valve wings; expanding keel single, stout, central on valve, divided at apex; covering membranes reduced to a limb or 0; closing bodies 0
Seeds highly variable in size, shape, colour and texture
x = 9
Flowering autumn to early winter
Distinguishing characters:
Compact dwarf perennials, mostly underground with paired leaves fused to form ± obconic bodies with flat or rounded, often windowed and ornamented tops
Fruit 4-7(9)-locular
Pedicels ebracteate
Nomenclature:
Lithops N.E.Br.
Brown: 44 (1922)
Friedrich: 68 (1970)
Herre: 196 (1971)
Cole (1988)
Clark: 1 (1996)
Smith et al.: 112 (1998)
Distribution & Notes:
Southern Africa: Species 36, widespread, Namibia, Botswana and South Africa, but excluding Mpumalanga and KwaZulu-Natal and occurs mainly in the Nama Karoo, succulent karoo and in rare cases in semi-arid savanna regions
References:
BROWN, N.E. 1922. Mesembryanthemum and some new genera separated from it. Gardeners' Chronicle: 71
CLARK, J.Y. 1996. A key to Lithops (Aizoaceae). Bradleya 14
COLE, D. 1988. Lithops. Flowering stones. Acorn Books in association with Russel Friedman Books, South Africa
FRIEDRICH, H.C. 1970. Aizoaceae. Prodromus einer Flora von Südwestafrika 27
HERRE, H. 1971. The genera of the Mesembryanthemaceae. Tafelberg, Cape Town
SMITH, G.F., CHESSELET, P., VAN JAARSVELD, E.J., HARTMANN, H., HAMMER, S., VAN WYK, B-.E., BURGOYNE, P., KLAK, C. & KURZWEIL, H. 1998. Mesembs of the world. Briza, Pretoria
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