Mesembryanthemaceae - Ruschioideae - Leipoldtia Group - Cephalophyllum N.E.Br.
Description:
Prostrate perennials, sometimes compact, rarely erect and shrubby, with a prominent, vegetative, monopodial head in several species; main stem short, producing decumbent or prostrate branches, with distinct internodes between some leaf pairs
Leaves opposite, not connate, crowded at end of stem and branches, apically triangular in cross section, basally rounded, rarely quilt-shaped, club-shaped or triquetrous, often mucronate; stomata not sunken
Flowers 1-3, rarely solitary, terminal, up to 50 mm in diameter, on elongate, bibracteate or ebracteate pedicels; opening midday or afternoon
Sepals 5
Petals free, linear, yellow, purple or white in various combinations
Stamens erect, often differing in colour from petals
Nectary a crenulate ring
Ovary with top raised into a crater-like rim surrounding stigmas or sometimes simply depressed in centre; placentas basal; stigmas 10-21, plumose
Fruit a 10-21-locular capsule, of Leipoldtia type; with a prominent closing bulge formed by roof of locule; valve wings well developed; expanding keels parallel; covering membranes present; closing bodies large; sometimes as tumble fruit
Seeds ovoid or pear-shaped, small, brown
x = 9 (polyploidy)
Flowering in winter
Distinguishing characters:
Cushion-forming perennials, creeping or erect
Flowers often brilliantly multicoloured
Fruit 10-20-locular with large closing bodies, broad valve wings and covering membranes with large closing bulges distally
Nomenclature:
Cephalophyllum N.E.Br.
Brown: 433 (1925)
Friedrich: 20 (1970)
Herre: 108 (1971)
Hartmann: 264 (1978)
Hartmann: 653 (1983)
Hartmann: 141 (1984)
Hartmann: 263 (1986)
Hartmann: 93 (1988)
Smith et al.: 256 (1998)
Distribution & Notes:
Southern Africa: Species 33, in a broad band from Lüderitz to Aus in Namibia, stretching southwards through the Northern Cape and into the Western Cape, occasionally reaching the sea, and extending as far east as Oudtshoorn
References:
BROWN, N.E. 1925. Mesembryanthemum and some new genera separated from it. Gardeners' Chronicle 78
FRIEDRICH, H.C. 1970. Aizoaceae. Prodromus einer Flora von Südwestafrika 27
HARTMANN, H.E.K. 1978. Zur Kenntnis der Gattung Cephalophyllum. Botanische Jahrbücher 99
HARTMANN, H.E.K. 1983. Interaction of ecology, taxonomy and distribution in some Mesembryanthemaceae. Bothalia 14
HARTMANN, H.E.K. 1984. Zur Biologie und Taxonomie des Cephalophyllum-curtophyllum-Komplexes. Mitteilungen aus dem Institut für Allgemeine Botanik. Hamburg 19
HARTMANN, H.E.K. 1986: Chromosome numbers in the genus Cephalophyllum N.E.Br. (Mesembryanthemaceae). Notes on Cephalophyllum V. Cactus and Succulent Journal (U.S) 58
HARTMANN, H.E.K. 1988. Monographien der Subtribus Leipoldtiinae. VIII. Monographie der Gattung Cephalophyllum (Mesembryanthemaceae). Mitteilungen aus dem Institut für Allgemeine Botanik. Hamburg 22
HERRE, H. 1971. The genera of the Mesembryanthemaceae. Tafelberg, Cape Town
SMITH, G.F., CHESSELET, P., VAN JAARSVELD, E.J., HARTMANN, H., HAMMER, S., VAN WYK, B-.E., BURGOYNE, P., KLAK, C. & KURZWEIL, H. 1998. Mesembs of the world. Briza, Pretoria
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