Trees or shrubs; plants monoecious or dioecious; leaves and shoot apices deciduous
Leaves distichously arranged, entire or lobed; 3-5-nerved, petiolate; stipules free, almost lateral, small, caducous
Flowers in solitary axillary, unisexual spikes
Male flowers in catkins; tepals membranous, 4-segmented, imbricate; stamens 4, filaments free, inflexed in bud
Female flowers in short, dense heads; tepals 4, decussate, imbricate, becoming succulent in fruit
Ovary enclosed within calyx, ovoid or subglobose; ovule pendulous; style short or obsolete; stigmas 2, (sub)equal
Fruit a ± succulent syncarp, of persistent fleshy tepals and exocarp; endocarp crustaceous
Seeds small, with endosperm; cotyledons thin, equal, plane, with membranous testa
x = 14 (can also be 7) (high polyploidy, B-chromosomes)
Nomenclature:
Morus L.
Linnaeus: 986 (1753)
Linnaeus: 424 (1754)
Adanson: 377 (1763)
Jussieu: 402 (1789)
Endlicher: 33 (1847)
Bentham: 364 (1880)
Engler: 72 (1889)
Engler: 3 (1898)
Rendle: 19 (1916)
Berg: 335 (1977)
Berg: 2 (1989)
Berg: 13 (1991)
Distribution & Notes:
Global: Species ± 13, in temperate to subtropical regions of the Old and New World
Southern Africa: Only 1 species native in Africa: Morus mesozygia Stapf, KwaZulu-Natal; 2 species are escapes from cultivation: *Morus alba L. (the White Mulberry) and *Morus japonica Audib.
References:
ADANSON, M. 1763. Famille des Chataigniers. Castaneae. Familles des plantes 2. Vincent, Paris
BENTHAM, G. 1880. Tribus Moreae. In G. Bentham & J.D. Hooker, Genera plantarum 3. Lovell Reeve & Co., London
BERG, C.C. 1977. Revisions of African Moraceae (excluding Dorstenia, Ficus, Masanga, and Myrianthus). Bulletin du Jardin Botanique National de Belgique 47
BERG, C.C. 1989. Flora of tropical East Africa. Moraceae
BERG, C.C. 1991. Moraceae. Flora zambesiaca 9,6
ENDLICHER, S.L. 1847. Moreae. Genera plantarum supplementum quartum. Beck, Vienna
ENGLER, A. 1889. Moraceae. Die natürlichen Pflanzenfamilien 3,1
ENGLER, A. 1898. Moraceae (excl. Ficus). Moraceae africanae. Monographieen afrikanischer Pflanzenfamilien und -gattungen. Vol. 1. Engelmann, Leipzig
JUSSIEU, A.L. DE. 1789. Urticeae. Genera plantarum secundum ordines naturales disposita. Herissant & Barrois, Paris
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754. Genera plantarum, edn 5. Laurentius Salvius, Stockholm
RENDLE, A.B. 1916. Moraceae. Morus & Cardiogyne. Flora of tropical Africa 6,2
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