Annual, biennial or perennial herbs, usually less than 0.3 m tall, often tufted or cushion-like
Leaves small, variable in shape, with cauline ones opposite, petiolate or sessile; stipules 0
Inflorescence often a panicle or cyme, terminal or rarely axillary, sometimes flowers solitary; bracts herbaceous
Flowers bisexual, subperigynous
Sepals 5, very rarely 4, free
Petals 5, white, cream or pink, entire or emarginate, rarely 0
Stamens 5 or 10, arising on a glandular perigynous disc
Ovary 1-locular; ovules many on a free-central placenta; styles (2)3(-5), episepalous
Fruit a capsule, subglobose, oblong or pyriform, dehiscent, mostly by twice the number of teeth as styles present, by 2 or 3(-5) bifid valves
Seeds many, compressed, smooth or tuberculate
x = 10, 11 (7, 8, 9, 12, 13) (high polyploidy)
Nomenclature:
*Arenaria L.
Linnaeus: 423 (1753)
Linnaeus: 193 (1754)
Jussieu: 301 (1789)
Candolle: 400 (1824)
Pax: 84 (1889)
Williams: 326 (1898)
Pax & Hoffmann: 315 (1934)
McNeill: 102 (1962)
McNeill: 245 (1963)
Chater & Halliday: 116 (1964)
Meikle: 261 (1977)
Bittrich: 225 (1993)
Distribution & Notes:
Global: Species ± 150, mainly in northern temperate regions, also in the arctic, some species on the mountains of South America and Northeast Africa
Southern Africa: Species 2 introduced, infrequent
References:
BITTRICH, V. 1993. Caryophyllaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
CANDOLLE, A.-P. DE. 1824. Caryophylleae. Prodromus 1. Treuttel & Würtz, Paris
JUSSIEU, A.L. DE. 1789. Caryophylleae, les caryophyllées. Genera plantarum secundum ordines naturales disposita. Herissant & Barrois, Paris
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754. Genera plantarum, edn 5. Laurentius Salvius, Stockholm
MCNEILL, J. 1962. Taxonomic studies in the Alsinoideae 1. Generic and infra-generic groups. Notes from the Royal Botanic Garden Edinburgh 24,2
MCNEILL, J. 1963. Taxonomic studies in the Alsinoideae II. A revision of the species in the Orient. Notes from the Royal Botanic Garden Edinburgh 24,3
MEIKLE, R.D. 1977. Caryophyllaceae. Flora of Cyprus 1
PAX, F. 1889. Caryophyllaceae. Die natürlichen Pflanzenfamilien 3,1b
PAX, F. & HOFFMANN, K. 1934. Caryophyllaceae. Die natürlichen Pflanzenfamilien II,16c
WILLIAMS, F.N. 1898. A revision of the genus Arenaria L. The Journal of the Linnean Society (Botany) 33
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