Erect, tall, annual, aromatic herbs without latex, dioecious, rarely monoecious; most parts of plant with minute adpressed swollen-based glandular hairs. Male and female plants dimorphic:
Males taller and more slender with longer narrower leaflets, inflorescence sparsely leafy, plant dying soon after flowering
Female shorter, more robust, inflorescence densely leafy, plant living several months after pollination
Leaves alternate (opposite at stem base), petiolate, palmately compound or lobed, leaflets uneven in size, serrate; stipules lateral, linear, acute, persistent
Male inflorescences axillary, much-branched, lax, cymose panicles, bristly-hairy, exceeding leaves but bearing few scattered leaves
Male flowers small, pedicellate, regular; perianth uniseriate, lobes 5, free, imbricate, greenish or whitish, boat-shaped, spreading or reflexed; stamens 5, opposite perianth lobes, erect in bud, pendulous at maturity, dehiscence longitudinal, basipetal; filaments short; pistillode 0
Female inflorescences axillary, short, compact, not exceeding leaves, few-flowered
Female flowers in pairs, each with stipule-like bract and small green organ ('bracteole' or 'calyx') completely enveloping ovary and loosely enclosing mature fruit, forming basally swollen tubular sheath acuminate at apex and covered with fine hairs and short-stalked or sessile resinous glands; perianth thin, undivided, tightly enveloping ovary and mature fruit (often reduced or absent in cultivated forms), marbled with light and dark areas; ovary superior, sessile, ± globose, 1-locular; ovule solitary, anatropous, bitegmic, crassinucellar and pendent from near apex of locule; style short; stigma branches 2, long, filiform, densely pubescent, caducous
Fruit a globular to ovoid achene tightly covered by thin crustaceous perianth, reticulate venation of surface beneath perianth visible
Southern Africa: Species 1: *Cannabis sativa L. with 3 varieties, 'Dagga', probably originally from Asia and now widely cultivated and naturalised worldwide in warmer areas
References:
ADANSON, M. 1763. Famille les Chataigners. Castaneae. Familles des plantes 2. Paris
ANDERSON, L.C. 1980. Leaf variation among Cannabis species from a controlled garden. Botanical Museum Leaflets Harvard University 28
BAILLON, H.E. 1875. Ulmacées. Histoire des plantes 6. Hachette & Co., Paris
BENTHAM, G. 1880. Urticaceae. Tribus Cannabineae. In G. Bentham & J.D. Hooker, Genera plantarum 3,1. Lovell Reeve & Co., London
BURTT DAVY, J. 1932. Cannabaceae. A manual of the flowering plants and ferns of the Transvaal with Swaziland 2. Longmans, Green & Co., London
CANDOLLE, A. DE. 1869. Cannabineae. In A. de Candolle, Prodromus 16,1. Masson & Sons, Paris
ENGLER, A. 1888. Moraceae. Subfamily Cannaboideae. Die natürlichen Pflanzenfamilien III,1
HENDERSON, M.D. & ANDERSON, J.G. 1966. Common weeds in South Africa. Memoirs of the Botanical Survey of South Africa No. 37
JUSSIEU, A.L. DE. 1789. Urticae. Genera plantarum secundum ordines naturales disposita. Treuttel & Würtz, Paris
KUBITZKI, K. 1993. Cannabaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
LAMARCK, J.B.A.P.M. DE. 1785. Chanvre, Cannabis. Dictionnaire encyclopédique de botanique 1, 2. H. Agasse, Paris
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754. Genera plantarum, edn 5. Laurentius Salvius, Stockholm
MILLER, N.G. 1970 (for a detailed account and extensive bibliography). The genera of the Cannabaceae in the southeastern United States. Journal of the Arnold Arboretum 51
RENDLE, A.B. 1916. Cannabinaceae. Flora of tropical Africa 6,2
SCHULTES, R.E., KLEIN, W.M., PLOWMAN, T. & LOCKWOOD, T.E. 1974. An example of taxonomic neglect. Botanical Museum Leaflets Harvard University 23
SMALL, E. 1997. Cannabaceae. Flora of North America north of Mexico 3
SMALL, E. & CRONQUIST, A. 1976. A practical and natural taxonomy for Cannabis. Taxon 25
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