Dwarf shrubs, dioecious, mostly many-stemmed and often much and intricately branched, with thick woody roots
Leaves decussate or (rarely) in whorls of 3 but often seemingly in much larger numbers at nodes, small and often ± ericoid, ± acute, acuminate to mucronate at apex; stipular sheaths (sub)sessile, small, ± cup-shaped, with or without 1(-3) minute setae on either side
Inflorescence frequently leafy and inconspicuous, made up of subsessile, mostly 3-1-flowered cymes, arranged in pairs, or single at nodes, or flowers solitary and terminal on shoots
Flowers subsessile, subtended by a pair of leafy bracts, 4- or 5(6)-merous
Calyx with lobes small or subobsolete
Male flowers: corolla with tube short, ± cylindrical to broadly funnel-shaped, lobes recurved, ± lanceolate; anthers yellowish to whitish, exserted, dangling on long, slender, filiform filaments; minute rudimentary ovary usually present
Female flowers: corolla much smaller; tube cylindrical, lobes linear to ± lanceolate, erect to ± spreading
Ovary 2-locular; ovule 1 per locule, erect, from base of locule; style 0; stigmas 2, long-exserted, hairy, often purplish red
Fruit crowned by persistent calyx lobes, sometimes supported by a small ± U-shaped carpophore, dehiscent, occasionally ± inflated, or hard, indehiscent, without carpophore
x = 11 (aneuploids)
Nomenclature:
Nenax Gaertn.
Gaertner: 165 (1788)
Hooker: 140 (1873)
Schumann: 129 (1891)
Phillips: 736 (1951)
Puff: 37 (1986)
Distribution & Notes:
Southern Africa: Species 9, endemic, S Namibia, North-West, Free State, Lesotho, Northern, Western and Eastern Cape
References:
GAERTNER, C.F. 1788. De fructibus et seminibus plantarum 1. G.H. Schramm, Tübingen
HOOKER, J.D. 1873. Rubiaceae. In G. Bentham & J.D. Hooker, Genera plantarum 2. L. Reeve, London
PHILLIPS, E.P. 1951. The genera of South African flowering plants. Memoirs of the Botanical Survey of South Africa No. 25
PUFF, C. 1986. Paederieae, Anthospermeae, Rubieae. Flora of southern Africa 31, part 1, fascicle 2
SCHUMANN, K.M. 1891. Rubiaceae. Die natürlichen Pflanzenfamilien 4,4
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