Trees or many-stemmed shrubs, dioecious or polygamous, rarely monoecious; bark often peeling or flaking, with resin ducts, secreting an odoriferous resin
Branchlets often spine-tipped
Leaves alternate, simple, trifoliolate or imparipinnate; stipules 0
Flowers unisexual, rarely bisexual, regular, usually small, perigynous to hypogynous; in axillary simple or compound dichasial or paniculate cymes, or singly in clusters; male flowers usually larger than female flowers
Calyx infundibuliform, campanulate with 4 valvate, persistent lobes
Petals 4, free or united near base, usually yellow to green
Disc in perigynous flowers adnate to hypanthium, cylindrical; in hypogynous flowers not adnate to calyx or corolla, intrastaminal, cylindrical, usually with 4 large or with 4 large and 4 small lobes
Stamens 8(4), obdiplostemonous, 4 antesepalous stamens longer than other 4, arising on outside or on top of disc; anthers introrse, with longitudinal slits; staminodes present in female flowers
Ovary half-inferior or superior; rudimentary in male flowers; 2-locular; with 2 epitropous ovules per locule, pendulous on axile placenta; style short; stigma capitate, obscurely 2-4-lobed
Fruit an ovoid to subglobose drupe; in ripe fruit the thin exocarp and fleshy mesocarp splitting into 2 longitudinal valves to expose a crustaceous or bony endocarp (putamen), usually clasped by a red or yellowish pseudo-aril; pseudo-aril either cupular with short lobes or arms, or with 2-4 relatively long arms, or covering ± whole putamen without distinct arms; putamen usually enclosing 1 fertile locule and 1 much smaller abortive locule
Seeds: embryo straight, cotyledons much folded
x = 13 (polyploidy)
Nomenclature:
Burseraceae
Wild: 263 (1963)
Merxmüller: 1 (1968)
Van der Walt: 53 (1973)
Van der Walt: 195 (1975)
Van der Walt: 5 (1986)
Distribution & Notes:
Global: Genera 16, with ± 500 species distributed mainly in the tropics, but especially in Malaysia, Africa and South America
Southern Africa: Genus 1, species 31
References:
MERXMÜLLER, H. 1968. Burseraceae. Prodromus einer Flora von Südwestafrika 70
VAN DER WALT, J.J.A. 1973. The South African species of Commiphora. Bothalia 11
VAN DER WALT, J.J.A. 1975. The South West African species of Commiphora. Mitteilungen der Botanischen Staatssammlung München 12
VAN DER WALT, J.J.A. 1986. Burseraceae. Flora of southern Africa 18, 3
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