Annual or perennial, erect or diffuse herbs, rarely woody at base, usually many-stemmed, sometimes viscid, raphides often present in all parts; stems slender, glabrous or pubescent, often with glandular rings about internodes
Leaves opposite, commonly in pairs of unequal size at each node, often fleshy, entire or repand, petiolate or sessile
Inflorescences terminal and axillary, paniculate, umbellate or cymose, sometimes diffuse, pedunculate; bracteoles 1 or 2, minute, caducous
Flowers bisexual, small, solitary or glomerate, subsessile to pedicellate, fugacious
Perianth: tube variable in length, lower portion constricted above ovary, cylindrical to obconical, upper portion petaloid, coloured, campanulate, caducous after anthesis, with lower part persistent and becoming hardened in fruit, sometimes glandular; lobes 5
Stamens 1-4(-6), exserted or included, unequal; filaments filiform, shortly connate below; anthers usually subglobular
Ovary ellipsoid, oblique, shortly stipitate; style filiform, exserted or included; stigma capitate or peltate
Fruit enclosed in lower portion of perianth (anthocarp); anthocarp fusiform, valvate or turbinate, conspicuously 3-5-sulcate, symmetrical, glabrous or glandular pubescent, often viscid; ribs rounded, acute or winged, usually mucous when wet
Seed with scanty endosperm; embryo hooked
x = 13 (polyploidy)
Nomenclature:
Boerhavia L.
Linnaeus: 3 (1753) 'Boerhaavia'
Linnaeus: 9 (1754)
Jussieu: 91 (1789)
Brown: 422 (1810)
Choisy: 449 (1849)
Hooker: 5 (1880)
Heimerl: 26 (1889)
Baker & Wright: 2 (1909)
Cooke: 393 (1910)
Standley: 204 (1918)
Burtt Davy: 209 (1926)
Heimerl: 117 (1934)
Stemmerik: 452 (1964) in part
Codd: 113 (1966)
Schreiber: 1 (1969)
Fosberg: 1 (1978)
Stirton: 79 (1982)
Stannard: 20 (1988)
Bittrich & Kühn: 481 (1993)
Thulin: 169 (1993)
Whitehouse: 2 (1996)
Distribution & Notes:
Global: Species ± 20, in almost all tropical and subtropical areas of the world, mostly in North America; several are cosmopolitan weeds
Southern Africa: Species 7, fairly widespread throughout warm, arid regions, often found in disturbed areas, Namibia, Botswana, Northern Province, Gauteng, Mpumalanga, Swaziland, Free State, KwaZulu-Natal, Northern, Western (only Beaufort-West District) and Eastern Cape
References:
BAKER, J.G. & WRIGHT, C.H. 1909. Nyctagineae. Flora of tropical Africa 6,1
BITTRICH, V. & KÜHN, U. 1993. Nyctaginaceae. In K. Kubitzki, J.G. Rohwer & V. Bittrich, The families and genera of vascular plants - dicotyledons 2. Springer-Verlag, Berlin
BROWN, R. 1810. Nyctagines Juss. Prodromus florae Novae Hollandiae et Insulae van-Diemen 1. Johnson & Co., London
BURTT DAVY, J. 1926. Nyctaginaceae. A manual of the flowering plants and ferns of the Transvaal with Swaziland 1. Longmans, Green & Co., London
CHOISY, J.D. 1849. Nyctagineae. In A. de Candolle, Prodromus 13,2. V. Masson, Paris
CODD, L.E.W. 1966. Notes on Boerhavia in southern Africa. Bothalia 9
COOKE, T. 1910. Nyctagineae. Flora capensis 5,1
FOSBERG, F.R. 1978. Studies in the genus Boerhavia L. (Nyctaginaceae). Smithsonian Contributions to Botany 39
HEIMERL, A. 1889. Nyctaginaceae. Die natürlichen Pflanzenfamilien 3,1b
HEIMERL, A. 1934. Nyctaginaceae. Die natürlichen Pflanzenfamilien, edn 2, 16c
HOOKER, J.D. 1880. Nyctaginaceae. In G. Bentham & J.D. Hooker, Genera plantarum 3,1. Lovell Reeve & Co., London
JUSSIEU, A.L. DE. 1789. Nyctagines, les Nictages. Genera plantarum secundum ordines naturales disposita. Herissant & Barrois, Paris
LINNAEUS, C. 1753. Species plantarum, edn 1. Laurentius Salvius, Stockholm
LINNAEUS, C. 1754. Genera plantarum, edn 5. Laurentius Salvius, Stockholm
SCHREIBER, A. 1969. Nyctaginaceae. Prodromus einer Flora von Südwestafrika 25
STIRTON, C.H. 1982. Nyctaginaceae. The identity of Boerhaavia pterocarpa in South Africa. Bothalia 14
THULIN, M. 1993. Nyctaginaceae. Flora of Somalia 1
WHITEHOUSE, C. 1996. Flora of tropical East Africa. Nyctaginaceae
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