Poaceae - Pooideae - Aveneae - Aveninae - Holcus L.
Description:
Annual or perennial, tufted
Leaf blade linear, expanded, pubescent or pilose; ligule an unfringed or fringed membrane
Inflorescence a panicle, contracted, often dense, sometimes almost spike-like, oblong or interrupted; spikelets solitary, pedicelled, pedicel terete, pilose
Spikelet laterally compressed, falling with glumes; glumes ± equal, ± as long as to longer than spikelet, membranous, dissimilar, keeled; lower glume lanceolate, hairy on keels and margins, 1-3-nerved, awned; awn half as long as body of glume; upper glume larger, elliptic-lanceolate or ovate, 3-nerved, with 2 prominent lateral nerves, hairy on keel and margins, shortly mucronate or awned; awn as long as body of glume
Florets 2, a brief rachilla extension often present; lower floret bisexual; lemma firmer in texture than glumes, shiny, ovate, rounded on back, obscurely 3-5-nerved, obtuse or acute, awnless; upper floret specialised in form, male or sterile, lemma shorter to ± as long as lower lemma, lanceolate, hyaline or membranous, glabrous, awned from back near apex; awn short, stout, hooked, usually geniculate, slightly twisted; palea smaller than lemma, narrow-lanceolate
Lodicules 2, delicate
Stamens 3
Ovary glabrous; styles distinct, plumose above
Caryopsis narrowly ellipsoid
x = 4, 7 (polyploidy)
Nomenclature:
Holcus L.
Linnaeus: 1047 (1753) name conserved
Stapf: 464 (1899)
Stent: 284 (1924)
Chippindall: 87 (1955)
Tutin: 230 (1980)
Clayton & Renvoize: 130 (1986)
Gibbs Russell et al.: 180 (1990)
Watson & Dallwitz: 464 (1994)
Distribution & Notes:
Global: Species 6, Europe, North Africa and Middle East
Southern Africa: Species ± 2, 1 endemic: Holcus setiger Nees, 1 naturalised: *H. lanatus L., northern provinces of South Africa, KwaZulu-Natal, Eastern and Western Cape
References:
CHIPPINDALL, L.K.A. 1955. A guide to the identification of grasses in South Africa. In D. Meredith, The grasses and pastures of South Africa. Central News Agency, Cape Town
CLAYTON, W.D. & RENVOIZE S.A. 1986. Genera graminum. Grasses of the world. Kew Bulletin. Additional series 13
GIBBS RUSSELL, G.E., WATSON, L., KOEKEMOER, M., SMOOK, L., BARKER, N.P., ANDERSON, H.M. & DALLWITZ. M.J. 1990. Grasses of southern Africa. Memoirs of the Botanical Survey of South Africa No. 58
LINNAEUS, C. 1753. Species plantarum. Laurentius Salvius, Stockholm
STAPF, O. 1898-1900. Gramineae. Flora capensis 7
STENT, S.M. 1924. South African Gramineae. Grasses of the Transvaal as represented in the National Herbarium. Bothalia 1
TUTIN, T.G. 1980. Flora europaea 5
WATSON, L. & DALLWITZ, M.J. 1994. The grass genera of the world, revised edn. CAB International, Oxon
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