Trees, shrubs, shrublets or woody climbers, very rarely subherbaceous; lepidote or with microscopic and sometimes macroscopic stalked glands
Leaves opposite, or sometimes in whorls of 3 or 4, or rarely alternate, usually petiolate (rarely subsessile), almost always entire; petiole sometimes persisting (especially in climbers) forming a ± hooked spine
Flowers bisexual, regular, or slightly irregular, 4- or 5-merous, in elongated or subcapitate axillary or extra-axillary spikes or racemes or in terminal or terminal and axillary, often leafy panicles
Receptacle usually clearly divided into a lower part and an upper part which varies from patelliform to infundibuliform and is itself sometimes visibly differentiated into a lower part containing the disc (when present) and an often more expanded upper part
Sepals: lobes 4 or 5 (rarely more), deltate to ± subulate or filiform, sometimes scarcely developed
Petals usually 4 or 5 (rarely absent, in aberrant specimens and up to 7 in occasional flowers), small and inconspicuous to showy and exceeding the sepals, of various colours
Stamens twice as many as petals, inserted in 1 or more, usually 2, series inside upper receptacle and usually exserted
Disc glabrous or hairy, with or without a free margin, sometimes inconspicuous or absent
Style free (in southern Africa species); stigma sometimes ± expanded
Ovary completely inferior
Fruit 4- or 5-winged, ridged or angled, sessile or stipitate, indehiscent or rarely tardily dehiscent; pericarp usually thin and papery, sometimes leathery, more rarely fleshy
Seeds: cotyledons various
x = 13 (11, 12, 14) (high polyploidy)
Nomenclature:
Combretum Loefl.
Loefling: 308 (1758) name conserved
Sonder: 508 (1862)
Exell: 101 (1978)
Van Wyk: 125 (1984)
Carr & Rogers: 173 (1987)
Carr: 13 (1988)
Carr & Retief: 38 (1989)
Rodman: 45 (1990)
Hennessy & Rodman: 149 (1995)
Poivrea Thouars
Thouars: 28 (1811)
Sonder: 512 (1862)
Distribution & Notes:
Global: Species 250, fairly cosmopolitan in warm climates, excluding Australia and Pacific Islands
Southern Africa: Species ± 30, widespread in all countries and provinces, except in Free State (rare), Lesotho and Western Cape
References:
CARR, J.D. 1988. Combretaceae in southern Africa. Tree Society of southern Africa, Johannesburg
CARR, J.D. & RETIEF, E. 1989. A new species of Combretum from Natal. Bothalia 19
CARR, J.D. & ROGERS, B.C. 1987. Chemosystematic studies of the genus Combretum (Combretaceae), part 1. South African Journal of Botany 53
HENNESSY, E.F. & RODMAN, S. 1995. A note on Combretum subgenus Combretum section Macrostigmatea (Combretaceae). Bothalia 25
LOEFLING, P. 1758. Iter Hispanicum. Lars Salvius Kostnad, Stockholm
RODMAN, S. 1990. The validity of currently recognised sectional limits within Combretum Loefling, subgenus Combretum (Combretaceae) in southern Africa. M.Sc. thesis, Department of Botany in Faculty of Science at the University of Durban-Westville, Durban
THOUARS, L.M.A.A. DU PETIT 1811. Observations sur les plantes des îles de France. (Mél. Bot. III). A. Bertrand, Paris
VAN WYK, A.E. 1984. A new species of Combretum from Venda and taxonomic notes on the section Angustimarginata (Combretaceae). South African Journal of Botany 3
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