Leaf blade linear, expanded or folded, sometimes setaceous; ligule a hyaline or scarious, unfringed membrane
Inflorescence a narrow panicle, contracted or open, erect or nodding; spikelets solitary, pedicelled
Spikelet large, 7-30 mm long, laterally compressed, disarticulating above glumes and between florets; glumes ± equal or unequal, usually shorter than spikelet, hyaline or subhyaline, acute or acuminate or shortly awned, ± keeled; lower glume 1-3-nerved; upper glume 3(-7)-nerved
Florets 2-6, bisexual; uppermost floret male, sterile or reduced; rachilla hairy, rarely glabrous, sometimes produced into a short bristle beyond uppermost floret; lemma firmer than glumes, firmly membranous to leathery with scarious or hyaline tips, glabrous, rounded on back, acute or acuminate, 5-11-nerved, 2-lobed (rarely 4-lobed), lobes awnless or awned; central awn from upper part of back between lobes, awn longer than body of lemma, geniculate and twisted, or slightly so; callus hairy; palea shorter than lemma, 2-keeled, ciliate
Lodicules 2, large, hyaline
Stamens 3
Ovary hairy from above middle or at apex only; styles distinct, short, plumose
x = 7 (high polyploidy)
Nomenclature:
Helictotrichon Besser
Besser: 526 ['326'] (1827)
Stapf: 472 (1899) under Avenastrum Jess.
Stent: 284 (1924) under Avenastrum Jess.
Schweickerdt: 185 (1937)
Phillips: 118 (1951)
Chippindall: 76 (1955)
Clayton: 86 (1970)
Sevenster & Veldkamp: 329 (1983)
Clayton & Renvoize: 123 (1986)
Gibbs Russell et al.: 175 (1990)
Watson & Dallwitz: 446 (1994)
Distribution & Notes:
Global: Species ± 100, mainly temperate Eurasia, but extending across the tropical mountains to temperate regions throughout the world
Southern Africa: Species ± 12, northern provinces of South Africa, Swaziland, Free State, KwaZulu-Natal, Lesotho, Western and Eastern Cape
References:
BESSER, W.S.J.T. VON 1827. Helictotrichon. In J.A. Schultes & J.H. Schultes, Mantissa 3. J.C. Cotta, Stuttgart
CHIPPINDALL, L.K.A. 1955. A guide to the identification of grasses in South Africa. In D. Meredith, The grasses and pastures of South Africa. Central News Agency, Cape Town
CLAYTON, W.D. 1970. Flora of tropical East Africa. Gramineae (Part 1)
CLAYTON, W.D. & RENVOIZE S.A. 1986. Genera graminum. Grasses of the world. Kew Bulletin. Additional series 13
GIBBS RUSSELL, G.E., WATSON, L., KOEKEMOER, M., SMOOK, L., BARKER, N.P., ANDERSON, H.M. & DALLWITZ. M.J. 1990. Grasses of southern Africa. Memoirs of the Botanical Survey of South Africa No. 58
PHILLIPS, E.P. 1951. The genera of South African flowering plants. Memoirs of the Botanical Survey of South Africa No. 25
SCHWEICKERDT, H.G. 1937. A revision of the South African species of Helictotrichon. Bothalia 3
SEVENSTER, J.C. & VELDKAMP, J.F. 1983. A revision of Helictotrichon (Gramineae) in Malasia. Blumea 28
STAPF, O. 1898-1900. Gramineae. Flora capensis 7
STENT, S.M. 1924. South African Gramineae. Grasses of the Transvaal as represented in the National Herbarium. Bothalia 1
WATSON, L. & DALLWITZ, M.J. 1994. The grass genera of the world, revised edn. CAB International, Oxon
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