Annual or perennial herbs or shrublets, generally erect, sometimes prostrate, usually woolly or cobwebby, often glandular, rarely glabrous
Leaves
alternate, straight, variously shaped, sessile or petiolate, often hairy, pilose to velutinous, margins entire, flat or sometimes revolute
Capitula
disciform or discoid; cylindric, turbinate, campanulate, subglobose or depressed-globose, in terminal, loose, flat-topped corymbs or solitary
Involucral bracts
in few to many rows; stereome usually fenestrated (divided); white, yellow, straw-coloured, brown, pink, purple, or red, sometimes a combination of colours; glabrous or hairy
Receptacle
smooth, honeycombed or fimbrilliferous, flat, epaleate, rarely paleate
Marginal florets
female, fewer or sometimes more than central florets; corolla often narrowly tubular, sometimes with well-developed lamina, rarely filiform, yellow
Central florets
bisexual; corolla funnel-shaped or tubular and then ± campanulate above, lobes 5, hairs always present on backs of lobes, yellow
Anthers
ecalcarate, caudate; with lanceolate apical appendages; endothecial tissue polarised
Style
branches truncate, penicillate; stigmatic areas separated
Cypselas
sparsely hairy with short, clavate, myxogenic hairs or with long nonmyxogenic hairs or glabrous; epidermis smooth or often papillose
Pappus
of barbellate or subplumose bristles, generally in 1 row; bristles basally with or without patent cilia, apical cells clavate or not
x = 7 (4, 5, 6, 8, 13) (aneuploids, high polyploidy)
Nomenclature:
Helichrysum
Mill.
Miller: [462] (1754)
Persoon: 414 (1807) name conserved
Hilliard: 61 (1983)
Anderberg: 141 (1991)
Anderberg: 358 (1994)
Achyrocline
(Less.) DC.
Hilliard & Burtt: 201 (1981)
Hilliard: 59 (1983)
Mesfin Tadesse & Reilly: 379 (1995).
Leontonyx
Cass.
Cassini: 466 (1822)
Distribution & Notes:
Global
: Species ± 600, largely Africa and Madagascar, also Europe, Asia and Australia
Southern Africa
: Species 244, widely distributed
References:
ANDERBERG, A.A. 1991. Taxonomy and phylogeny of the tribe
Gnaphalieae
(
Asteraceae
).
Opera Botanica
104
ANDERBERG, A.A. 1994. Tribes
Inuleae
,
Plucheeae
,
Gnaphalieae
. In K. Bremer,
Asteraceae
, cladistics and classification
. Timber Press, Portland, Oregon
CASSINI, A.H.G. DE. 1822.
Dictionnaire des sciences naturelles
25. Ed. F. Cuvier. edn 2. Le Normant, Paris
HILLIARD, O.M. & BURTT, B.L. 1981. Some generic concepts in
Compositae
-
Gnaphaliinae
.
Botanical Journal of the Linnean Society
82
MESFIN TADESSE & REILLY, T. 1995. 17. A contribution to studies on
Helichrysum
(
Compositae
-
Gnaphalieae
) - a revision of the species of north-east tropical Africa. In D.J.H. Hind, C. Jeffrey & G.V. Pope,
Advances in Compositae Systematics
. Royal Botanic Gardens, Kew
MILLER, P. 1754.
The gardener's dictionary
(Abridged edition), edn 4. Rivington, London
PERSOON, C.H. 1807.
Helichrysum
.
Synopsis plantarum
. 2. Cramer, Paris
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