Stems
arising from subterranean, thick woody tubers or rhizomes
Leaves
alternate or crowded on brachyblasts, sessile, straight, adaxially concave, glabrous above, glandular-hairy below, margin entire
Capitula
radiate, solitary, on pedunculoid few-leaved stems or on long leafless peduncles
Involucral bracts
with spathulate laminas
Receptacle
flat or flat to convex, epaleate
Ray florets
female, yellow with purple bands dorsally, in one row, fewer than disc florets
Cypselas
as in disc florets
Pappus
of narrow scales
Disc florets
bisexual, corolla yellow
Anthers
ecalcarate, with short tails; endothecial tissue polarised; apical appendage acute
Style
branches truncate with obtuse sweeping hairs apically
Cypselas
narrowly elliptic with 5 or 3 vascular bundles, glabrous or sparsely hairy with elongated twin hairs
Pappus
of 5 barbellate capillary bristles and narrow scales, or of ± connate scales only
Nomenclature:
Comborhiza
Anderb. & K.Bremer
Anderberg & Bremer: 1070 (1991)
Anderberg: 330 (1994)
Distribution & Notes:
Southern Africa
: Species 2, endemic,
Comborhiza longipes
(K.Bremer) Anderb. & K.Bremer, Western Cape, and
C. virgata
(N.E.Br.) Anderb. & K.Bremer, KwaZulu-Natal
References:
ANDERBERG, A.A. 1994. Tribes
Inuleae
,
Plucheeae
,
Gnaphalieae
. In K. Bremer,
Asteraceae
, cladistics and classification
. Timber Press, Portland, Oregon
ANDERBERG, A.A. & BREMER, K. 1991. Parsimony analysis and cladistic reclassification of the
Relhania
generic group (
Asteraceae
-
Gnaphalieae
).
Annals of the Missouri Botanical Garden
78
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