Perennial herbs or dwarf shrublets; stems leafy throughout, pubescent, hairs retrorse and minutely glandular
Leaves opposite below, alternate above, simple, variously elliptical and oblong to oblanceolate, often narrowed to petiole-like, sometimes half-clasping or connate base, usually toothed, thick or leathery, glandular-punctate, sometimes also with glandular and/or eglandular hairs; radical leaves sometimes present
Flowers in a terminal, cone-shaped, very congested head, nodding in flower, erect and elongated in fruit
Bracts broad, clearly distinct from leaves, adnate to pedicel, free from calyx
Calyx unequally 5-lobed, rarely 2 anterior lobes obsolete, split nearly or quite to base at least on anterior side, thin-textured
Corolla slightly bilabiate, glabrous, thick, leathery when dry; tube funnel-shaped; posterior lobes exterior in bud, lobes spreading, oblong to suborbicular, entire, margins pale, thin-textured
Stamens 4, didynamous, arising near top of corolla tube, 2 or 4 exserted; posterior filaments decurrent ± to base of tube; anthers synthecate, dorsifixed; staminodes 0
Nectary at base of ovary on posterior side
Ovary bilocular, ± oblong; ovules 7-many per locule; stigma lingulate, flattened, with 2 marginal bands of stigmatic papillae, exserted
Fruit a septicidal capsule with a short loculicidal split at tip of each valve, glabrous
Seeds angled by pressure, sinuously colliculate, reddish brown
x = 7
Nomenclature:
Glumicalyx Hiern
Hiern: t. 2769 (1903)
Hiern: 369 (1904)
Marloth: 129 (1932)
Junell: 172 (1961)
Hilliard & Burtt: 155 (1977)
Hilliard: 526 (1994)
Distribution & Notes:
Southern Africa: Species 6, mainly Lesotho, also nearby in South Africa, in Free State, KwaZulu-Natal and Eastern Cape, along Drakensberg from Harrismith and Golden Gate to Barkly East and Lady Grey
HILLIARD, O.M. & BURTT, B.L. 1977. Notes on some plants of southern Africa chiefly from Natal: VI. Notes from the Royal Botanic Garden Edinburgh 35
HILLIARD, O.M. 1994. The Manuleae. A tribe of Scrophulariaceae. Edinburgh University Press, Edinburgh
JUNELL, S. 1961. Ovarian morphology and taxonomical position of Selagineae. Svensk Botanisk Tidskrift 55
MARLOTH, R. 1932. Scrophulariaceae. The flora of South Africa 3(1). Darter Bros. & Co., Cape Town
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