Scrophulariaceae - Scrophularioideae - Gratioleae - Limosella L.
Description:
Marsh or aquatic, perennial herbs, tufted, creeping or floating; stemless or with stolons, rooting at nodes, glabrous
Leaves usually radical, sometimes fasciculate at nodes, rarely alternate, simple, usually petiolate, erect or floating, lamina linear, subulate, spathulate, oblong-ovate or suborbicular
Flowers usually solitary, axillary; pedicels ebracteolate, often deflexed in fruit
Calyx tubular, (4)5-lobed; tube funnel-shaped or campanulate; lobes subequal, usually shorter than tube, rarely longer
Corolla tubular, 5-lobed, subregular; tube campanulate or turbinate; limb erect or rotate; lobes subequal, posterior outside in bud, rounded, ovate or oblong, pilose on inner face
Stamens 4, mostly somewhat didynamous, in mouth of corolla tube or shortly exserted; filaments filiform, arising ± in middle of corolla tube, filaments of lower stamens often crossing those immediately above them at right angles; anthers with 2 equal, confluent thecae
Ovary bilocular, at least near base, globose or elliptical; ovules many; style varying in length, terete, sometimes incurved, included or exserted; stigma capitate
Fruit an ovoid to ± spherical capsule, almost indehiscent or at length loculicidal; valves entire
Seeds many, subglobose, 4-sided, or oblong; testa thin, brown, ribbed or reticulate
x = 10 (9 - 1 report) (polyploidy)
Nomenclature:
Limosella L.
Linnaeus: 631 (1753)
Hiern: 356 (1904)
Glück: 488 (1934)
Merxmüller & Roessler: 26 (1967)
Hilliard & Burtt: 212 (1986)
Philcox: 73 (1990)
Mielcarek: 64 (1996)
Distribution & Notes:
Global: Species ± 11, cosmopolitan, mainly in temperate and cooler regions but also in tropical mountains
Southern Africa: Species ± 8, widespread but not in Swaziland
References:
GLÜCK, H. 1934. Beiträge zur Systematik, Morphologie und Biologie der Gattung Limosella. Botanische Jahrbücher 66
HILLIARD, O.M. & BURTT, B.L. 1986. Notes on some plants of southern Africa chiefly from Natal: XII. Notes from the Royal Botanic Garden Edinburgh 43
LINNAEUS, C. 1753. Species plantarum, edn 1. Laurentius Salvius, Stockholm
MERXMÜLLER, H. & ROESSLER, H. 1967. Scrophulariaceae. Prodromus einer Flora von Südwestafrika 126
MIELCAREK, R. 1996. Les Scrophulariaceae dans la flore d'Afrique centrale (excl. Linderneae). Fragmenta floristica et geobotanica 41
PHILCOX, D. 1990. Scrophulariaceae. Flora zambesiaca 8,2
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