Annual or perennial herbs or undershrubs, sprawling to erect; stems often quadrangular and branched from base, glabrous or pubescent
Leaves mostly opposite, sometimes alternate above, linear or lanceolate to ovate, entire or toothed, sessile or petiolate
Flowers solitary-axillary or in terminal racemes, bracteate, pedicellate, often alternate
Bracts sometimes leafy
Calyx 5-lobed, somewhat accrescent in fruit
Corolla bilabiate, personate, 5-lobed; tube short, extended at front into a dependent spur, pocket or pouch; posterior lip 4-lobed, exterior in bud, with laterals outside both posteriormost lobes and anterior lip; anterior lip 1-lobed, entire or emarginate, with a convex, sometimes papillate palate at its base
Stamens 4, didynamous, arising in mouth of corolla tube; filaments linear, anterior pair usually bent at base round posterior pair and coming to be in posterior position, usually appendaged at base; anthers usually cohering in pairs about stigma; thecae confluent
Ovary bilocular, usually elliptical; ovules many; style short, thick and terete or filiform; stigma simple or bilobed
Fruit a septicidal capsule, usually ovate to urceolate, somewhat laterally compressed; apex rounded to emarginate; valves boat-shaped
Seeds many in 1 or 2 series, usually ± ellipsoid, more rarely spindle-shaped; wing membranous, entire or interrupted
x = 9 (10 - 1 report, aneuploids)
Nomenclature:
Nemesia Vent.
Ventenat: 41, t. 41 (I804)
Hiern: 169 (1904)
Diels: 121 (1910)
Merxmüller & Roessler: 34 (1967)
Hilliard & Burtt: 385 (1986)
Steiner: 405 (1989)
Philcox: 9 (1990)
Steiner: 63 (1996)
Distribution & Notes:
Global: Species ± 65, Africa; tropical (few) and southern Africa
Southern Africa: Species ± 60, in all provinces but mostly in Western Cape
References:
DIELS, L. 1910. Formationen und Florenelemente im nordwestlichen Kapland. Scrophulariaceae. Botanische Jahrbücher 44
HILLIARD, O.M. & BURTT, B.L. 1986. Notes on some plants of southern Africa chiefly from Natal: XIII. Notes from the Royal Botanic Garden Edinburgh 43
MERXMÜLLER, H. & ROESSLER, H. 1967. Scrophulariaceae. Prodromus einer Flora von Südwestafrika 126
PHILCOX, D. 1990. Scrophulariaceae. Flora zambesiaca 8,2
STEINER, K.E. 1989. A new perennial Nemesia (Scrophulariaceae) from the western Cape. South African Journal of Botany 55
STEINER, K.E. 1996. Chromosome numbers and relationships in tribe Hemimerideae (Scrophulariaceae). Systematic Botany 21
VENTENAT, É.P. 1804. Jardin de la Malmaison 1. Crapelet (typ.), Paris
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