Annual herbs, erect, prostrate, decumbent or ascending, or floating, mostly glabrous, sometimes pubescent or subglabrous, often glandular-punctate; stems simple or branching, often rooting at nodes
Leaves opposite, petiolate to sessile, variously obovate to spathulate or linear to lanceolate or ovate, entire or variously dentate to crenate, or leaves capillary-divided in aquatic species; often glandular punctate
Flowers axillary, usually solitary, sometimes several, pedicellate, usually bibracteolate just below calyx
Calyx 5-partite; sepals imbricate in bud, unequal, posterior broadest, laterals usually very narrow
Corolla bilabiate; tube cylindrical; upper lip emarginate or 2-lobed, exterior in bud; lower lip 3-lobed; lobes equal or subequal
Stamens 4, didynamous, perfect, included, arising in corolla tube; anthers bithecate; thecae usually parallel, sometimes divaricate or finally confluent; staminodes 0
Nectary 0 or 1
Ovary bilocular; ovules many; style dilated at apex, entire, concave or 2-lobed
Fruit an ovoid to globose or ellipsoid capsule, loculicidal or septicidal, bivalved; valves bifid or bipartite
Seeds small, many, usually striate and transversely reticulate
x = 8 (1 report, polyploidy)
Nomenclature:
Bacopa Aubl.
Aublet: 128, t. 49 (1775) name conserved
Hepper: 358 (1963)
Philcox: 48 (1990)
Greuter et al.: 298 (1994)
Mielcarek: 33 (1996)
Moniera P.Browne
Browne: 269 (1756)
Hiern: 355 (1904)
Distribution & Notes:
Global: Species ± 56, tropical and subtropical regions
Southern Africa: Species 3, Botswana, Northern Province, Mpumalanga and KwaZulu-Natal; marsh plants, at times aquatic
References:
AUBLET, J.B.C.F. 1775. Bacopa. Histoire des plantes de la Guiane Française 1. Didot, Paris
BROWNE, P. 1756. Moniera. The civil and natural history of Jamaica in three parts. Published by the author, London
GREUTER, W., BARRIE, F.R., BURDET, H.M., CHALONER, W.G., DEMOULIN, V., HAWKSWORTH, D.L., JØRGENSEN, P.M., NICOLSON, D.H., SILVA, P.C., TREHANE, P, & MCNEILL, J. 1994. Appendix IIIA, Nomina generica conservanda et rejicienda. International code of botanical nomenclature. Koeltz Scientific Books, Königstein
HEPPER, F.N. 1963. Scrophulariaceae. Flora of west tropical Africa, edn 2, 2
MIELCAREK, R. 1996. Les Scrophulariaceae dans la flore d'Afrique centrale (excl. Linderneae). Fragmenta floristica et geobotanica 41
PHILCOX, D. 1990. Scrophulariaceae. Flora zambesiaca 8,2
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